Rps26 (ribosomal protein S26) - Rat Genome Database

Send us a Message
Submit Data |  Help |  Video Tutorials |  News |  Publications |  Download |  REST API |  Citing RGD |  Contact   
Search RGD Grant Resources Citing RGD About Us Contact Us
Genes Variants Community Projects QTLs Strains Markers Genome Information Ontologies Cell Lines References Download Submit Data
OntoMate (Literature Search) JBrowse (Genome Browser) Synteny Browser (VCMap) Variant Visualizer Multi-Ontology Enrichment (MOET) Gene-Ortholog Location Finder (GOLF) InterViewer (Protein-Protein Interactions) PhenoMiner (Quatitative Phenotypes) Gene Annotator OLGA (Gene List Generator) AllianceMine GViewer (Genome Viewer)
Aging & Age-Related Disease Behavioral & Substance Use Disorder Cancer & Neoplastic Disease Cardiovascular Disease Coronavirus Disease Developmental Disease Diabetes Hematologic Disease Immune & Inflammatory Disease Infectious Disease Liver Disease Neurological Disease Obesity & Metabolic Syndrome Renal Disease Respiratory Disease Sensory Organ Disease
Find Models Genetic Models Autism Models Rat PhenoMiner (Quantitative Phenotypes) Chinchilla PhenoMiner Expected Ranges (Quantitative Phenotype) PhenoMiner Term Comparison Hybrid Rat Diversity Panel Phenotypes Phenotypes in Other Animal Models Animal Husbandry Strain Medical Records Phylogenetics Strain Availability Calendar Rats 101 Submissions Photo Archive
Pathways
Rat Community Forum Directory of Rat Laboratories Video Tutorials News RGD Publications RGD Presentations Archive Nomenclature Guidelines Resource Links Laboratory Resources Employment Resources
Advanced Search (OLGA)
Gene: Rps26 (ribosomal protein S26) Rattus norvegicus
Analyze
Symbol: Rps26
Name: ribosomal protein S26
RGD ID: 621044
Description: Predicted to enable mRNA binding activity. Predicted to be a structural constituent of ribosome. Predicted to be involved in cytoplasmic translation and negative regulation of RNA splicing. Part of cytosolic small ribosomal subunit. Human ortholog(s) of this gene implicated in Diamond-Blackfan anemia 10. Orthologous to human RPS26 (ribosomal protein S26); PARTICIPATES IN ribosome biogenesis pathway; translation pathway; INTERACTS WITH 2,3,7,8-tetrachlorodibenzodioxine; ammonium chloride; bisphenol A.
Type: protein-coding
RefSeq Status: PROVISIONAL
Previously known as: 40S ribosomal protein S26; small ribosomal subunit protein eS26
RGD Orthologs
Human
Mouse
Black Rat
Alliance Orthologs
More Info more info ...
Related Pseudogenes: LOC108349054   Rps26-ps11   Rps26l5  
Latest Assembly: GRCr8 - GRCr8 Assembly
NCBI Annotation Information: Annotation category: partial on reference assembly
Position:
Rat AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
GRCr871,641,846 - 1,643,404 (-)NCBIGRCr8GRCr8GRCr8
GRCr8 Ensembl71,641,421 - 1,643,399 (-)EnsemblGRCr8
mRatBN7.271,057,332 - 1,058,882 (-)NCBImRatBN7.2mRatBN7.2
mRatBN7.2 EnsemblX107,822,178 - 107,822,525 (-)EnsemblmRatBN7.2
UTH_Rnor_SHR_Utx73,820,252 - 3,821,824 (-)NCBIUTH_Rnor_SHR_UtxUTH_Rnor_SHR_Utx
UTH_Rnor_SHRSP_BbbUtx_1.075,696,245 - 5,697,817 (-)NCBIUTH_Rnor_SHRSP_BbbUtx_1.0UTH_Rnor_SHRSP_BbbUtx_1.0
UTH_Rnor_WKY_Bbb_1.075,992,952 - 5,994,529 (-)NCBIUTH_Rnor_WKY_Bbb_1.0UTH_Rnor_WKY_Bbb_1.0
Rnor_6.073,052,112 - 3,053,662 (-)NCBIRnor_6.0Rnor_6.0rn6Rnor6.0
Rnor_5.073,025,307 - 3,070,860 (-)NCBIRnor_5.0Rnor_5.0rn5
RGSC_v3.471,919,153 - 1,920,703 (-)NCBIRGSC_v3.4RGSC_v3.4rn4
Celera7927,684 - 929,234 (-)NCBICelera
RGSC_v3.171,919,152 - 1,920,703 (-)NCBI
Cytogenetic Map7q11NCBI
JBrowse:




Gene-Chemical Interaction Annotations     Click to see Annotation Detail View
(+)-catechin  (ISO)
(1->4)-beta-D-glucan  (ISO)
1,2-dimethylhydrazine  (ISO)
17beta-estradiol  (ISO)
17beta-hydroxy-17-methylestra-4,9,11-trien-3-one  (ISO)
2,2',4,4'-Tetrabromodiphenyl ether  (ISO)
2,3,7,8-tetrachlorodibenzodioxine  (EXP,ISO)
2,4,6-tribromophenol  (ISO)
2,6-dimethoxyphenol  (ISO)
3,3',5,5'-tetrabromobisphenol A  (ISO)
4,4'-diaminodiphenylmethane  (ISO)
4,4'-sulfonyldiphenol  (ISO)
5-fluorouracil  (ISO)
actinomycin D  (ISO)
ammonium chloride  (EXP)
arsane  (ISO)
arsenic atom  (ISO)
arsenite(3-)  (ISO)
benzo[a]pyrene  (ISO)
bis(2-ethylhexyl) phthalate  (ISO)
bisphenol A  (EXP,ISO)
bisphenol AF  (ISO)
Bisphenol B  (ISO)
bromobenzene  (EXP)
cadmium atom  (ISO)
cadmium dichloride  (EXP,ISO)
caffeine  (ISO)
chlorpyrifos  (ISO)
choline  (ISO)
chromium(6+)  (ISO)
copper atom  (EXP)
copper(0)  (EXP)
copper(II) sulfate  (ISO)
Cuprizon  (EXP)
DDE  (ISO)
decabromodiphenyl ether  (ISO)
diquat  (ISO)
diuron  (ISO)
enzyme inhibitor  (ISO)
epoxiconazole  (ISO)
folic acid  (ISO)
furfural  (ISO)
glafenine  (EXP)
glyphosate  (ISO)
isoniazide  (ISO)
isoprenaline  (ISO)
ivermectin  (ISO)
L-methionine  (ISO)
lead(0)  (ISO)
leflunomide  (ISO)
lipopolysaccharide  (ISO)
manganese atom  (ISO)
manganese(0)  (ISO)
manganese(II) chloride  (ISO)
methylparaben  (ISO)
miconazole  (ISO)
nickel atom  (ISO)
nitric oxide  (ISO)
Nutlin-3  (ISO)
perfluorooctane-1-sulfonic acid  (ISO)
phenobarbital  (ISO)
quercetin  (ISO)
S-butyl-DL-homocysteine (S,R)-sulfoximine  (ISO)
sodium arsenite  (ISO)
sodium chloride  (ISO)
testosterone  (ISO)
tetrachloromethane  (EXP)
triphenyl phosphate  (ISO)
triptonide  (ISO)
valproic acid  (ISO)

Gene Ontology Annotations     Click to see Annotation Detail View

Biological Process

Cellular Component

Molecular Function

Molecular Pathway Annotations     Click to see Annotation Detail View
References

References - curated
# Reference Title Reference Citation
1. Structures of the human and Drosophila 80S ribosome. Anger AM, etal., Nature. 2013 May 2;497(7447):80-5. doi: 10.1038/nature12104.
2. Isolation of eukaryotic ribosomal proteins. Purification and characterization of the 40 S ribosomal subunit proteins Sa, Sc, S3a, S3b, S5', S9, S10, S11, S12, S14, S15, S15', S16, S17, S18, S19, S20, S21, S26, S27', and S29. Collatz E, etal., J Biol Chem. 1977 Dec 25;252(24):9071-80.
3. Phylogenetic-based propagation of functional annotations within the Gene Ontology consortium. Gaudet P, etal., Brief Bioinform. 2011 Sep;12(5):449-62. doi: 10.1093/bib/bbr042. Epub 2011 Aug 27.
4. Molecular cloning and nucleotide sequence of DNA complementary to rat ribosomal protein S26 messenger RNA. Kuwano Y, etal., J Biochem (Tokyo) 1985 Apr;97(4):983-92.
5. Electronic Transfer of LocusLink and RefSeq Data NCBI rat LocusLink and RefSeq merged data July 26, 2002
6. Functional dichotomy of ribosomal proteins during the synthesis of mammalian 40S ribosomal subunits. O'Donohue MF, etal., J Cell Biol. 2010 Sep 6;190(5):853-66. doi: 10.1083/jcb.201005117.
7. OMIM Disease Annotation Pipeline OMIM Disease Annotation Pipeline
8. GOA pipeline RGD automated data pipeline
9. ClinVar Automated Import and Annotation Pipeline RGD automated import pipeline for ClinVar variants, variant-to-disease annotations and gene-to-disease annotations
10. Data Import for Chemical-Gene Interactions RGD automated import pipeline for gene-chemical interactions
11. Comprehensive gene review and curation RGD comprehensive gene curation
12. Affinity labelling of rat liver ribosomal protein S26 by heptauridylate containing a 5'-terminal alkylating group. Stahl J and Kobetz ND, Mol Biol Rep. 1984 Jan;9(4):219-22.
13. Generation and initial analysis of more than 15,000 full-length human and mouse cDNA sequences. Strausberg RL, etal., Proc Natl Acad Sci U S A. 2002 Dec 24;99(26):16899-903. Epub 2002 Dec 11.
Additional References at PubMed
PMID:15489334   PMID:25002582   PMID:25763846   PMID:29476059  


Genomics

Comparative Map Data
Rps26
(Rattus norvegicus - Norway rat)
Rat AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
GRCr871,641,846 - 1,643,404 (-)NCBIGRCr8GRCr8GRCr8
GRCr8 Ensembl71,641,421 - 1,643,399 (-)EnsemblGRCr8
mRatBN7.271,057,332 - 1,058,882 (-)NCBImRatBN7.2mRatBN7.2
mRatBN7.2 EnsemblX107,822,178 - 107,822,525 (-)EnsemblmRatBN7.2
UTH_Rnor_SHR_Utx73,820,252 - 3,821,824 (-)NCBIUTH_Rnor_SHR_UtxUTH_Rnor_SHR_Utx
UTH_Rnor_SHRSP_BbbUtx_1.075,696,245 - 5,697,817 (-)NCBIUTH_Rnor_SHRSP_BbbUtx_1.0UTH_Rnor_SHRSP_BbbUtx_1.0
UTH_Rnor_WKY_Bbb_1.075,992,952 - 5,994,529 (-)NCBIUTH_Rnor_WKY_Bbb_1.0UTH_Rnor_WKY_Bbb_1.0
Rnor_6.073,052,112 - 3,053,662 (-)NCBIRnor_6.0Rnor_6.0rn6Rnor6.0
Rnor_5.073,025,307 - 3,070,860 (-)NCBIRnor_5.0Rnor_5.0rn5
RGSC_v3.471,919,153 - 1,920,703 (-)NCBIRGSC_v3.4RGSC_v3.4rn4
Celera7927,684 - 929,234 (-)NCBICelera
RGSC_v3.171,919,152 - 1,920,703 (-)NCBI
Cytogenetic Map7q11NCBI
RPS26
(Homo sapiens - human)
Human AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
GRCh381256,041,918 - 56,044,697 (+)NCBIGRCh38GRCh38hg38GRCh38
GRCh38.p14 Ensembl1256,041,351 - 56,044,697 (+)Ensemblhg38GRCh38
GRCh371256,435,702 - 56,438,481 (+)NCBIGRCh37GRCh37hg19GRCh37
Build 361254,721,953 - 54,724,274 (+)NCBIBuild 36Build 36hg18NCBI36
Build 341254,722,192 - 54,724,272NCBI
Celera1256,088,373 - 56,090,694 (+)NCBICelera
Cytogenetic Map12q13.2NCBI
HuRef1253,474,591 - 53,476,912 (+)NCBIHuRef
CHM1_11256,402,791 - 56,405,112 (+)NCBICHM1_1
T2T-CHM13v2.01256,008,421 - 56,011,764 (+)NCBIT2T-CHM13v2.0
Rps26
(Mus musculus - house mouse)
Mouse AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
GRCm3910128,460,398 - 128,462,375 (-)NCBIGRCm39GRCm39mm39
GRCm39 Ensembl10128,460,403 - 128,462,616 (-)EnsemblGRCm39 EnsemblGRCm39
GRCm3810128,624,529 - 128,626,506 (-)NCBIGRCm38GRCm38mm10GRCm38
GRCm38.p6 Ensembl10128,624,534 - 128,626,747 (-)Ensemblmm10GRCm38
MGSCv3710128,061,585 - 128,063,562 (-)NCBIMGSCv37MGSCv37mm9NCBIm37
MGSCv3610128,027,485 - 128,029,432 (-)NCBIMGSCv36mm8
Celera10131,016,529 - 131,018,506 (-)NCBICelera
Cytogenetic Map10D3NCBI
cM Map1077.11NCBI
Rps26
(Rattus rattus - black rat)
Black Rat AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
Rrattus_CSIRO_v11156,100,395 - 156,101,999 (-)NCBIRrattus_CSIRO_v1

Variants

.
Variants in Rps26
33 total Variants
miRNA Target Status (No longer updated)

Predicted Target Of
Summary Value
Count of predictions:104
Count of miRNA genes:78
Interacting mature miRNAs:79
Transcripts:ENSRNOT00000007304, ENSRNOT00000051134
Prediction methods:Microtar, Miranda, Rnahybrid
Result types:miRGate_prediction

The detailed report is available here: Full Report CSV TAB Printer

miRNA Target Status data imported from miRGate (http://mirgate.bioinfo.cnio.es/).
For more information about miRGate, see PMID:25858286 or access the full paper here.


QTLs in Region (GRCr8)
The following QTLs overlap with this region.    Full Report CSV TAB Printer Gviewer
RGD IDSymbolNameLODP ValueTraitSub TraitChrStartStopSpecies
1300176Hrtrt10Heart rate QTL 103.19heart pumping trait (VT:2000009)heart rate (CMO:0000002)7123748937Rat
2298550Neuinf6Neuroinflammation QTL 63.3nervous system integrity trait (VT:0010566)spinal cord RT1-B protein level (CMO:0002132)7129716167Rat
2290372Gluco33Glucose level QTL 332.71blood glucose amount (VT:0000188)blood glucose level area under curve (AUC) (CMO:0000350)7131777942Rat
1300149Cm6Cardiac mass QTL 64.09heart mass (VT:0007028)heart left ventricle weight to body weight ratio (CMO:0000530)71104117804Rat
634336Anxrr17Anxiety related response QTL 173.66locomotor behavior trait (VT:0001392)number of entries into a discrete space in an experimental apparatus (CMO:0000960)71509236116977875Rat
724560Plsm3Polydactyly-luxate syndrome (PLS) morphotypes QTL 30.0003tibia length (VT:0004357)tibia length (CMO:0000450)7134650782Rat
9590142Scort5Serum corticosterone level QTL 524.40.001blood corticosterone amount (VT:0005345)plasma corticosterone level (CMO:0001173)7132612984Rat
1582260Bw72Body weight QTL 723.20.0043body mass (VT:0001259)body weight (CMO:0000012)7139960528Rat
1582261Bw69Body weight QTL 693.20.0048body mass (VT:0001259)body weight (CMO:0000012)7139960528Rat
1582262Bw75Body weight QTL 7530.0038body mass (VT:0001259)body weight (CMO:0000012)7139960528Rat
7411566Bw136Body weight QTL 13610.40.001body mass (VT:0001259)body weight gain (CMO:0000420)7132612984Rat


Expression

RNA-SEQ Expression

alimentary part of gastrointestinal system
appendage
circulatory system
ectoderm
hemolymphoid system
mesenchyme
mesoderm
musculoskeletal system
nervous system
renal system
reproductive system
sensory system
visual system
7 1 18 52 67 138 95 11 51 36 32 17 17

Sequence

Nucleotide Sequences
RefSeq Transcripts NM_013224 (Get FASTA)   NCBI Sequence Viewer   Search GEO for Microarray Profiles
  XM_063263070 (Get FASTA)   NCBI Sequence Viewer   Search GEO for Microarray Profiles
GenBank Nucleotide AC128207 (Get FASTA)   NCBI Sequence Viewer   Search GEO for Microarray Profiles
  BC061561 (Get FASTA)   NCBI Sequence Viewer   Search GEO for Microarray Profiles
  CH474104 (Get FASTA)   NCBI Sequence Viewer   Search GEO for Microarray Profiles
  FQ210888 (Get FASTA)   NCBI Sequence Viewer   Search GEO for Microarray Profiles
  FQ211456 (Get FASTA)   NCBI Sequence Viewer   Search GEO for Microarray Profiles
  FQ216976 (Get FASTA)   NCBI Sequence Viewer   Search GEO for Microarray Profiles
  FQ217039 (Get FASTA)   NCBI Sequence Viewer   Search GEO for Microarray Profiles
  FQ217630 (Get FASTA)   NCBI Sequence Viewer   Search GEO for Microarray Profiles
  FQ220214 (Get FASTA)   NCBI Sequence Viewer   Search GEO for Microarray Profiles
  FQ220682 (Get FASTA)   NCBI Sequence Viewer   Search GEO for Microarray Profiles
  FQ220803 (Get FASTA)   NCBI Sequence Viewer   Search GEO for Microarray Profiles
  FQ221742 (Get FASTA)   NCBI Sequence Viewer   Search GEO for Microarray Profiles
  FQ221917 (Get FASTA)   NCBI Sequence Viewer   Search GEO for Microarray Profiles
  FQ222007 (Get FASTA)   NCBI Sequence Viewer   Search GEO for Microarray Profiles
  FQ222170 (Get FASTA)   NCBI Sequence Viewer   Search GEO for Microarray Profiles
  FQ222423 (Get FASTA)   NCBI Sequence Viewer   Search GEO for Microarray Profiles
  FQ222638 (Get FASTA)   NCBI Sequence Viewer   Search GEO for Microarray Profiles
  FQ223096 (Get FASTA)   NCBI Sequence Viewer   Search GEO for Microarray Profiles
  FQ223202 (Get FASTA)   NCBI Sequence Viewer   Search GEO for Microarray Profiles
  FQ223253 (Get FASTA)   NCBI Sequence Viewer   Search GEO for Microarray Profiles
  FQ223633 (Get FASTA)   NCBI Sequence Viewer   Search GEO for Microarray Profiles
  FQ224810 (Get FASTA)   NCBI Sequence Viewer   Search GEO for Microarray Profiles
  FQ229027 (Get FASTA)   NCBI Sequence Viewer   Search GEO for Microarray Profiles
  FQ233862 (Get FASTA)   NCBI Sequence Viewer   Search GEO for Microarray Profiles
  FQ233965 (Get FASTA)   NCBI Sequence Viewer   Search GEO for Microarray Profiles
  FQ234974 (Get FASTA)   NCBI Sequence Viewer   Search GEO for Microarray Profiles
  FQ235169 (Get FASTA)   NCBI Sequence Viewer   Search GEO for Microarray Profiles
  FQ235253 (Get FASTA)   NCBI Sequence Viewer   Search GEO for Microarray Profiles
  JAXUCZ010000007 (Get FASTA)   NCBI Sequence Viewer   Search GEO for Microarray Profiles
  X02414 (Get FASTA)   NCBI Sequence Viewer   Search GEO for Microarray Profiles

Ensembl Acc Id: ENSRNOT00000168392   ⟹   ENSRNOP00000104557
Type: CODING
Position:
Rat AssemblyChrPosition (strand)Source
GRCr8 Ensembl71,641,421 - 1,643,399 (-)Ensembl
RefSeq Acc Id: NM_013224   ⟹   NP_037356
RefSeq Status: PROVISIONAL
Type: CODING
Position:
Rat AssemblyChrPosition (strand)Source
GRCr871,641,854 - 1,643,404 (-)NCBI
mRatBN7.271,057,332 - 1,058,882 (-)NCBI
Rnor_6.073,052,112 - 3,053,662 (-)NCBI
Rnor_5.073,025,307 - 3,070,860 (-)NCBI
RGSC_v3.471,919,153 - 1,920,703 (-)RGD
Celera7927,684 - 929,234 (-)RGD
Sequence:
RefSeq Acc Id: XM_063263070   ⟹   XP_063119140
Type: CODING
Position:
Rat AssemblyChrPosition (strand)Source
GRCr871,641,846 - 1,643,194 (-)NCBI
Protein Sequences
Protein RefSeqs NP_037356 (Get FASTA)   NCBI Sequence Viewer  
  XP_063119140 (Get FASTA)   NCBI Sequence Viewer  
GenBank Protein AAH61561 (Get FASTA)   NCBI Sequence Viewer  
  CAA26264 (Get FASTA)   NCBI Sequence Viewer  
  EDL84832 (Get FASTA)   NCBI Sequence Viewer  
Ensembl Protein ENSRNOP00000104557
GenBank Protein P62856 (Get FASTA)   NCBI Sequence Viewer  
RefSeq Acc Id: NP_037356   ⟸   NM_013224
- UniProtKB: P62856 (UniProtKB/Swiss-Prot),   A6KSG0 (UniProtKB/TrEMBL),   D3Z8D7 (UniProtKB/TrEMBL)
- Sequence:
RefSeq Acc Id: XP_063119140   ⟸   XM_063263070
- Peptide Label: isoform X1
- UniProtKB: D3Z8D7 (UniProtKB/TrEMBL)
Ensembl Acc Id: ENSRNOP00000104557   ⟸   ENSRNOT00000168392

Protein Structures
Name Modeler Protein Id AA Range Protein Structure
AF-P62856-F1-model_v2 AlphaFold P62856 1-115 view protein structure

Transcriptome

eQTL   View at Phenogen
WGCNA   View at Phenogen
Tissue/Strain Expression   View at Phenogen


Additional Information

Database Acc Id Source(s)
AGR Gene RGD:621044 AgrOrtholog
BioCyc Gene G2FUF-35347 BioCyc
Ensembl Genes ENSRNOG00000029512 Ensembl
  ENSRNOG00000081095 Ensembl, ENTREZGENE
Ensembl Transcript ENSRNOT00000168392 ENTREZGENE
Gene3D-CATH 3.30.1740.20 UniProtKB/Swiss-Prot
IMAGE_CLONE IMAGE:6921494 IMAGE-MGC_LOAD
InterPro Ribosomal_eS26_CS UniProtKB/Swiss-Prot
  Ribosomal_S26e UniProtKB/Swiss-Prot
  Ribosomal_S26e_sf UniProtKB/Swiss-Prot
KEGG Report rno:27139 UniProtKB/Swiss-Prot
MGC_CLONE MGC:72903 IMAGE-MGC_LOAD
NCBI Gene 27139 ENTREZGENE
PANTHER 40S RIBOSOMAL PROTEIN S26-RELATED UniProtKB/Swiss-Prot
  PTHR12538 UniProtKB/Swiss-Prot
Pfam Ribosomal_S26e UniProtKB/Swiss-Prot
PhenoGen Rps26 PhenoGen
PROSITE RIBOSOMAL_S26E UniProtKB/Swiss-Prot
RatGTEx ENSRNOG00000029512 RatGTEx
  ENSRNOG00000081095 RatGTEx
UniProt A6KSG0 ENTREZGENE, UniProtKB/TrEMBL
  D3Z8D7 ENTREZGENE, UniProtKB/TrEMBL
  P62856 ENTREZGENE, UniProtKB/Swiss-Prot
UniProt Secondary P02383 UniProtKB/Swiss-Prot
  P70394 UniProtKB/Swiss-Prot
  Q06722 UniProtKB/Swiss-Prot


Nomenclature History
Date Current Symbol Current Name Previous Symbol Previous Name Description Reference Status
2004-09-10 Rps26  ribosomal protein S26      Symbol and Name status set to approved 1299863 APPROVED
2002-08-07 Rps26  ribosomal protein S26      Symbol and Name status set to provisional 70820 PROVISIONAL

RGD Curation Notes
Note Type Note Reference
gene_protein 115 amino acids with a molecular weight of 13,015 634033