Nptxr (neuronal pentraxin receptor) - Rat Genome Database

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Gene: Nptxr (neuronal pentraxin receptor) Rattus norvegicus
Analyze
Symbol: Nptxr
Name: neuronal pentraxin receptor
RGD ID: 628898
Description: Enables pentraxin receptor activity. Involved in response to hydrogen peroxide. Located in extracellular space. Biomarker of brain ischemia and congestive heart failure. Orthologous to human NPTXR (neuronal pentraxin receptor); INTERACTS WITH 2,3,7,8-tetrachlorodibenzodioxine; 3,3',4,4',5-pentachlorobiphenyl; 6-propyl-2-thiouracil.
Type: protein-coding
RefSeq Status: REVIEWED
Previously known as: Npr
RGD Orthologs
Human
Mouse
Chinchilla
Bonobo
Dog
Squirrel
Pig
Green Monkey
Naked Mole-Rat
Alliance Orthologs
More Info more info ...
Latest Assembly: GRCr8 - GRCr8 Assembly
Position:
Rat AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
GRCr87113,217,046 - 113,235,122 (-)NCBIGRCr8GRCr8GRCr8
GRCr8 Ensembl7113,217,050 - 113,235,156 (-)EnsemblGRCr8
mRatBN7.27111,336,660 - 111,354,737 (-)NCBImRatBN7.2mRatBN7.2
mRatBN7.2 Ensembl7111,336,664 - 111,354,802 (-)EnsemblmRatBN7.2
UTH_Rnor_SHR_Utx7113,084,519 - 113,102,580 (-)NCBIUTH_Rnor_SHR_UtxUTH_Rnor_SHR_Utx
UTH_Rnor_SHRSP_BbbUtx_1.07115,308,059 - 115,326,122 (-)NCBIUTH_Rnor_SHRSP_BbbUtx_1.0UTH_Rnor_SHRSP_BbbUtx_1.0
UTH_Rnor_WKY_Bbb_1.07115,277,412 - 115,295,475 (-)NCBIUTH_Rnor_WKY_Bbb_1.0UTH_Rnor_WKY_Bbb_1.0
Rnor_6.07121,011,678 - 121,029,754 (-)NCBIRnor_6.0Rnor_6.0rn6Rnor6.0
Rnor_6.0 Ensembl7121,011,680 - 121,029,754 (-)Ensemblrn6Rnor6.0
Rnor_5.07121,002,637 - 121,020,713 (-)NCBIRnor_5.0Rnor_5.0rn5
RGSC_v3.47118,023,653 - 118,041,729 (-)NCBIRGSC_v3.4RGSC_v3.4rn4
Celera7107,667,311 - 107,685,387 (-)NCBICelera
RGSC_v3.17118,058,154 - 118,075,959 (-)NCBI
Cytogenetic Map7q34NCBI
JBrowse:




Disease Annotations     Click to see Annotation Detail View

Gene Ontology Annotations     Click to see Annotation Detail View

Cellular Component

Molecular Function

References

References - curated
# Reference Title Reference Citation
1. Microarray analysis of oxidative stress regulated genes in mesencephalic dopaminergic neuronal cells: relevance to oxidative damage in Parkinson's disease. Anantharam V, etal., Neurochem Int. 2007 May;50(6):834-47. Epub 2007 Feb 23.
2. Regulation of neuronal type genes in congestive heart failure rats. Andersson KB, etal., Acta Physiol (Oxf). 2006 Jan;186(1):17-27.
3. Neuronal pentraxin receptor, a novel putative integral membrane pentraxin that interacts with neuronal pentraxin 1 and 2 and taipoxin-associated calcium-binding protein 49. Dodds DC, etal., J Biol Chem 1997 Aug 22;272(34):21488-94.
4. Phylogenetic-based propagation of functional annotations within the Gene Ontology consortium. Gaudet P, etal., Brief Bioinform. 2011 Sep;12(5):449-62. doi: 10.1093/bib/bbr042. Epub 2011 Aug 27.
5. DNA array reveals altered gene expression in response to focal cerebral ischemia. Kim YD, etal., Brain Res Bull. 2002 Sep 15;58(5):491-8.
6. Biochemical interactions of the neuronal pentraxins. Neuronal pentraxin (NP) receptor binds to taipoxin and taipoxin-associated calcium-binding protein 49 via NP1 and NP2. Kirkpatrick LL, etal., J Biol Chem. 2000 Jun 9;275(23):17786-92.
7. Presynaptic Neuronal Pentraxin Receptor Organizes Excitatory and Inhibitory Synapses. Lee SJ, etal., J Neurosci. 2017 Feb 1;37(5):1062-1080. doi: 10.1523/JNEUROSCI.2768-16.2016. Epub 2016 Dec 16.
8. Rat ISS GO annotations from MGI mouse gene data--August 2006 MGD data from the GO Consortium
9. GOA pipeline RGD automated data pipeline
10. ClinVar Automated Import and Annotation Pipeline RGD automated import pipeline for ClinVar variants, variant-to-disease annotations and gene-to-disease annotations
11. Data Import for Chemical-Gene Interactions RGD automated import pipeline for gene-chemical interactions
Additional References at PubMed
PMID:8889548   PMID:15673668   PMID:25449907  


Genomics

Comparative Map Data
Nptxr
(Rattus norvegicus - Norway rat)
Rat AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
GRCr87113,217,046 - 113,235,122 (-)NCBIGRCr8GRCr8GRCr8
GRCr8 Ensembl7113,217,050 - 113,235,156 (-)EnsemblGRCr8
mRatBN7.27111,336,660 - 111,354,737 (-)NCBImRatBN7.2mRatBN7.2
mRatBN7.2 Ensembl7111,336,664 - 111,354,802 (-)EnsemblmRatBN7.2
UTH_Rnor_SHR_Utx7113,084,519 - 113,102,580 (-)NCBIUTH_Rnor_SHR_UtxUTH_Rnor_SHR_Utx
UTH_Rnor_SHRSP_BbbUtx_1.07115,308,059 - 115,326,122 (-)NCBIUTH_Rnor_SHRSP_BbbUtx_1.0UTH_Rnor_SHRSP_BbbUtx_1.0
UTH_Rnor_WKY_Bbb_1.07115,277,412 - 115,295,475 (-)NCBIUTH_Rnor_WKY_Bbb_1.0UTH_Rnor_WKY_Bbb_1.0
Rnor_6.07121,011,678 - 121,029,754 (-)NCBIRnor_6.0Rnor_6.0rn6Rnor6.0
Rnor_6.0 Ensembl7121,011,680 - 121,029,754 (-)Ensemblrn6Rnor6.0
Rnor_5.07121,002,637 - 121,020,713 (-)NCBIRnor_5.0Rnor_5.0rn5
RGSC_v3.47118,023,653 - 118,041,729 (-)NCBIRGSC_v3.4RGSC_v3.4rn4
Celera7107,667,311 - 107,685,387 (-)NCBICelera
RGSC_v3.17118,058,154 - 118,075,959 (-)NCBI
Cytogenetic Map7q34NCBI
NPTXR
(Homo sapiens - human)
Human AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
GRCh382238,818,452 - 38,844,028 (-)NCBIGRCh38GRCh38hg38GRCh38
GRCh38.p14 Ensembl2238,818,452 - 38,844,028 (-)Ensemblhg38GRCh38
GRCh372239,214,457 - 39,240,033 (-)NCBIGRCh37GRCh37hg19GRCh37
Build 362237,544,403 - 37,569,963 (-)NCBIBuild 36Build 36hg18NCBI36
Celera2223,021,120 - 23,046,674 (-)NCBICelera
Cytogenetic Map22q13.1NCBI
HuRef2222,180,084 - 22,228,226 (-)NCBIHuRef
CHM1_12239,173,407 - 39,198,951 (-)NCBICHM1_1
T2T-CHM13v2.02239,288,799 - 39,314,406 (-)NCBIT2T-CHM13v2.0
Nptxr
(Mus musculus - house mouse)
Mouse AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
GRCm391579,670,552 - 79,688,910 (-)NCBIGRCm39GRCm39mm39
GRCm39 Ensembl1579,670,552 - 79,688,967 (-)EnsemblGRCm39 EnsemblGRCm39
GRCm381579,786,351 - 79,804,709 (-)NCBIGRCm38GRCm38mm10GRCm38
GRCm38.p6 Ensembl1579,786,351 - 79,804,766 (-)Ensemblmm10GRCm38
MGSCv371579,616,781 - 79,635,139 (-)NCBIMGSCv37MGSCv37mm9NCBIm37
MGSCv361579,613,606 - 79,631,964 (-)NCBIMGSCv36mm8
Celera1581,905,537 - 81,923,864 (-)NCBICelera
Cytogenetic Map15E1NCBI
cM Map1537.85NCBI
Nptxr
(Chinchilla lanigera - long-tailed chinchilla)
Chinchilla AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
ChiLan1.0 EnsemblNW_00495541324,925,784 - 24,942,292 (-)Ensembl
ChiLan1.0NW_00495541324,924,464 - 24,941,417 (-)NCBIChiLan1.0ChiLan1.0
NPTXR
(Pan paniscus - bonobo/pygmy chimpanzee)
Bonobo AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
NHGRI_mPanPan1-v22348,672,073 - 48,697,746 (-)NCBINHGRI_mPanPan1-v2
NHGRI_mPanPan12251,364,213 - 51,408,232 (-)NCBINHGRI_mPanPan1
Mhudiblu_PPA_v02219,731,443 - 19,757,125 (-)NCBIMhudiblu_PPA_v0Mhudiblu_PPA_v0panPan3
PanPan1.12237,548,222 - 37,562,249 (-)NCBIPanPan1.1PanPan1.1panPan2
NPTXR
(Canis lupus familiaris - dog)
Dog AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
CanFam3.11025,983,249 - 26,012,169 (+)NCBICanFam3.1CanFam3.1canFam3CanFam3.1
CanFam3.1 Ensembl1025,988,906 - 26,008,258 (+)EnsemblcanFam3CanFam3.1
Dog10K_Boxer_Tasha1025,944,929 - 25,967,924 (+)NCBIDog10K_Boxer_TashaDog10K_Boxer_Tasha
ROS_Cfam_1.01026,779,304 - 26,803,082 (+)NCBIROS_Cfam_1.0ROS_Cfam_1.0
ROS_Cfam_1.0 Ensembl1026,779,409 - 26,803,065 (+)EnsemblROS_Cfam_1.0 Ensembl
UMICH_Zoey_3.11026,501,288 - 26,525,031 (+)NCBIUMICH_Zoey_3.1
UNSW_CanFamBas_1.01026,812,143 - 26,835,887 (+)NCBIUNSW_CanFamBas_1.0
UU_Cfam_GSD_1.01026,988,882 - 27,012,645 (+)NCBIUU_Cfam_GSD_1.0UU_Cfam_GSD_1.0
Nptxr
(Ictidomys tridecemlineatus - thirteen-lined ground squirrel)
Squirrel AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
HiC_Itri_2NW_0244049459,538,163 - 9,557,786 (+)NCBIHiC_Itri_2
SpeTri2.0 EnsemblNW_0049364922,522,170 - 2,564,874 (+)EnsemblSpeTri2.0 Ensembl
SpeTri2.0NW_0049364922,546,774 - 2,568,206 (+)NCBISpeTri2.0SpeTri2.0SpeTri2.0
NPTXR
(Sus scrofa - pig)
Pig AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
Sscrofa11.1 Ensembl59,207,642 - 9,230,355 (+)EnsemblsusScr11Sscrofa11.1
Sscrofa11.159,207,295 - 9,230,354 (+)NCBISscrofa11.1Sscrofa11.1susScr11Sscrofa11.1
Sscrofa10.256,452,962 - 6,475,686 (+)NCBISscrofa10.2Sscrofa10.2susScr3
NPTXR
(Chlorocebus sabaeus - green monkey)
Green Monkey AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
ChlSab1.11921,481,245 - 21,507,680 (-)NCBIChlSab1.1ChlSab1.1chlSab2
Vero_WHO_p1.0NW_023666045104,235,945 - 104,277,243 (+)NCBIVero_WHO_p1.0Vero_WHO_p1.0
Nptxr
(Heterocephalus glaber - naked mole-rat)
Naked Mole-Rat AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
HetGla_female_1.0 EnsemblNW_0046247529,315,853 - 9,336,043 (+)EnsemblHetGla_female_1.0 EnsemblhetGla2HetGla_female_1.0 Ensembl
HetGla 1.0NW_0046247529,315,834 - 9,335,205 (+)NCBIHetGla 1.0HetGla 1.0hetGla2

Variants

.
Variants in Nptxr
56 total Variants
miRNA Target Status (No longer updated)

Predicted Target Of
Summary Value
Count of predictions:356
Count of miRNA genes:216
Interacting mature miRNAs:246
Transcripts:ENSRNOT00000004703
Prediction methods:Miranda, Targetscan
Result types:miRGate_prediction

The detailed report is available here: Full Report CSV TAB Printer

miRNA Target Status data imported from miRGate (http://mirgate.bioinfo.cnio.es/).
For more information about miRGate, see PMID:25858286 or access the full paper here.


QTLs in Region (GRCr8)
The following QTLs overlap with this region.    Full Report CSV TAB Printer Gviewer
RGD IDSymbolNameLODP ValueTraitSub TraitChrStartStopSpecies
1300179Kidm5Kidney mass QTL 53.51kidney mass (VT:0002707)left kidney wet weight (CMO:0000083)792388565119280177Rat
1331728Bp214Blood pressure QTL 2142.825arterial blood pressure trait (VT:2000000)mean arterial blood pressure (CMO:0000009)782111245123048330Rat
1331731Bp216Blood pressure QTL 2162.851arterial blood pressure trait (VT:2000000)mean arterial blood pressure (CMO:0000009)7104186212135371468Rat
1357338Stl17Serum triglyceride level QTL 173.23blood triglyceride amount (VT:0002644)serum triglyceride level (CMO:0000360)771621154114982716Rat
70159Bp61Blood pressure QTL 610.0001arterial blood pressure trait (VT:2000000)systolic blood pressure (CMO:0000004)773618702118618702Rat
631687Hcas1Hepatocarcinoma susceptibility QTL 13.90.001liver integrity trait (VT:0010547)liver tumorous lesion volume to total liver volume ratio (CMO:0001082)793302009131686183Rat
61397Bw17Body weight QTL 176.3body mass (VT:0001259)body weight (CMO:0000012)795485047137014596Rat
631504Cm27Cardiac mass QTL 273.45heart left ventricle mass (VT:0007031)heart left ventricle wet weight (CMO:0000071)746307490127866166Rat
634322Bw12Body weight QTL 120body mass (VT:0001259)body weight (CMO:0000012)785043242130043242Rat
70173Niddm19Non-insulin dependent diabetes mellitus QTL 194.330.00005blood glucose amount (VT:0000188)blood glucose level area under curve (AUC) (CMO:0000350)785497398130497398Rat
1549899Stresp8Stress response QTL 84.370.0008stress-related behavior trait (VT:0010451)defensive burying duration (CMO:0001961)792362341137014596Rat
2317052Aia17Adjuvant induced arthritis QTL 172.13joint integrity trait (VT:0010548)left rear ankle joint diameter (CMO:0002149)783626689128626689Rat
631529Tls2T-lymphoma susceptibility QTL 200.001thymus integrity trait (VT:0010555)percentage of study population developing T-cell lymphomas during a period of time (CMO:0001911)782111382132099989Rat
731176Glom5Glomerulus QTL 52.50.0035kidney glomerulus morphology trait (VT:0005325)count of superficial glomeruli not directly contacting the kidney surface (CMO:0001002)798549867137014596Rat
7411607Foco15Food consumption QTL 150.001eating behavior trait (VT:0001431)feed conversion ratio (CMO:0001312)777807807122807807Rat
631201Panci1Pancreas inflammation QTL 100.001pancreas integrity trait (VT:0010560)percentage of study population displaying chronic pancreatitis at a point in time (CMO:0001214)773557047118557047Rat
634336Anxrr17Anxiety related response QTL 173.66locomotor behavior trait (VT:0001392)number of entries into a discrete space in an experimental apparatus (CMO:0000960)71509236116977875Rat
1331768Kidm10Kidney mass QTL 104.62096kidney mass (VT:0002707)left kidney wet weight (CMO:0000083)782111245123048330Rat
61357Bp38Blood pressure QTL 381.60.052arterial blood pressure trait (VT:2000000)systolic blood pressure (CMO:0000004)740540077123682549Rat
2313102Bmd79Bone mineral density QTL 792.30.0001tibia mineral mass (VT:1000283)total volumetric bone mineral density (CMO:0001728)796700404118618702Rat
2316947Rf58Renal function QTL 587.8kidney glomerulus morphology trait (VT:0005325)index of glomerular damage (CMO:0001135)749682832115766396Rat
1331748Bp215Blood pressure QTL 2154.043arterial blood pressure trait (VT:2000000)mean arterial blood pressure (CMO:0000009)7110725024135371468Rat
10450862Kidm55Kidney mass QTL 554.9kidney mass (VT:0002707)kidney weight (CMO:0000081)772725037117725037Rat
61428Scl3Serum cholesterol level QTL 33.2blood cholesterol amount (VT:0000180)serum total cholesterol level (CMO:0000363)7100548246137014596Rat
1331746Kidm9Kidney mass QTL 93.934kidney mass (VT:0002707)right kidney wet weight (CMO:0000082)782111245134948181Rat
10450856Livw4Liver weight QTL 43.8liver mass (VT:0003402)liver weight (CMO:0000092)772725037117725037Rat
7411654Foco25Food consumption QTL 259.30.001eating behavior trait (VT:0001431)feed conversion ratio (CMO:0001312)777807807122807807Rat
2316955Stl24Serum triglyceride level QTL 247.1blood triglyceride amount (VT:0002644)plasma triglyceride level (CMO:0000548)749682832115766396Rat
10402855Bp379Blood pressure QTL 3790.21arterial blood pressure trait (VT:2000000)mean arterial blood pressure (CMO:0000009)778682549123682549Rat
2316952Pur22Proteinuria QTL 225.2urine protein amount (VT:0005160)urine protein level (CMO:0000591)749682832115766396Rat
631540Bw9Body weight QTL 94.5body mass (VT:0001259)body weight (CMO:0000012)771621154119349044Rat
1358891Bp265Blood pressure QTL 2652.21arterial blood pressure trait (VT:2000000)mean arterial blood pressure (CMO:0000009)771827901136544683Rat

Markers in Region
RH130990  
Rat AssemblyChrPosition (strand)SourceJBrowse
mRatBN7.27111,336,728 - 111,337,919 (+)MAPPERmRatBN7.2
Rnor_6.07121,011,747 - 121,012,937NCBIRnor6.0
Rnor_5.07121,002,706 - 121,003,896UniSTSRnor5.0
RGSC_v3.47118,023,722 - 118,024,912UniSTSRGSC3.4
Celera7107,667,380 - 107,668,570UniSTS
RH 3.4 Map7858.1UniSTS
Cytogenetic Map7q34UniSTS


Expression

RNA-SEQ Expression

alimentary part of gastrointestinal system
appendage
circulatory system
ectoderm
endocrine system
endoderm
exocrine system
hemolymphoid system
hepatobiliary system
integumental system
mesenchyme
mesoderm
musculoskeletal system
nervous system
renal system
reproductive system
respiratory system
sensory system
visual system
16 12 67 162 89 88 57 72 57 6 333 172 11 141 80 92 31 17 17

Sequence


Ensembl Acc Id: ENSRNOT00000004703   ⟹   ENSRNOP00000004703
Type: CODING
Position:
Rat AssemblyChrPosition (strand)Source
GRCr8 Ensembl7113,217,050 - 113,235,156 (-)Ensembl
mRatBN7.2 Ensembl7111,336,664 - 111,354,802 (-)Ensembl
Rnor_6.0 Ensembl7121,011,680 - 121,029,754 (-)Ensembl
RefSeq Acc Id: NM_030841   ⟹   NP_110468
RefSeq Status: REVIEWED
Type: CODING
Position:
Rat AssemblyChrPosition (strand)Source
GRCr87113,217,046 - 113,235,122 (-)NCBI
mRatBN7.27111,336,660 - 111,354,737 (-)NCBI
Rnor_6.07121,011,678 - 121,029,754 (-)NCBI
Rnor_5.07121,002,637 - 121,020,713 (-)NCBI
RGSC_v3.47118,023,653 - 118,041,729 (-)RGD
Celera7107,667,311 - 107,685,387 (-)RGD
Sequence:
Protein Sequences
Protein RefSeqs NP_110468 (Get FASTA)   NCBI Sequence Viewer  
GenBank Protein EDM15780 (Get FASTA)   NCBI Sequence Viewer  
Ensembl Protein ENSRNOP00000004703
GenBank Protein O35764 (Get FASTA)   NCBI Sequence Viewer  
RefSeq Acc Id: NP_110468   ⟸   NM_030841
- Peptide Label: precursor
- UniProtKB: O35764 (UniProtKB/Swiss-Prot)
- Sequence:
Ensembl Acc Id: ENSRNOP00000004703   ⟸   ENSRNOT00000004703
Protein Domains
Pentraxin (PTX)

Protein Structures
Name Modeler Protein Id AA Range Protein Structure
AF-O35764-F1-model_v2 AlphaFold O35764 1-494 view protein structure

Transcriptome

eQTL   View at Phenogen
WGCNA   View at Phenogen
Tissue/Strain Expression   View at Phenogen

Promoters
RGD ID:13695488
Promoter ID:EPDNEW_R6013
Type:single initiation site
Name:Nptxr_1
Description:neuronal pentraxin receptor
SO ACC ID:SO:0000170
Source:EPDNEW (Eukaryotic Promoter Database, http://epd.vital-it.ch/)
Experiment Methods:Single-end sequencing.
Position:
Rat AssemblyChrPosition (strand)Source
Rnor_6.07121,029,781 - 121,029,841EPDNEW

Additional Information

Database Acc Id Source(s)
AGR Gene RGD:628898 AgrOrtholog
BioCyc Gene G2FUF-32743 BioCyc
Ensembl Genes ENSRNOG00000016156 Ensembl, ENTREZGENE
Ensembl Transcript ENSRNOT00000004703 ENTREZGENE
Gene3D-CATH 2.60.120.200 UniProtKB/Swiss-Prot
InterPro ConA-like_dom_sf UniProtKB/Swiss-Prot
  Neuronal_Pentraxin_Related UniProtKB/Swiss-Prot
  Pentraxin-related UniProtKB/Swiss-Prot
NCBI Gene 81005 ENTREZGENE
PANTHER NEURONAL PENTRAXIN RECEPTOR UniProtKB/Swiss-Prot
  PENTRAXIN UniProtKB/Swiss-Prot
Pfam Pentaxin UniProtKB/Swiss-Prot
PhenoGen Nptxr PhenoGen
PRINTS PENTAXIN UniProtKB/Swiss-Prot
PROSITE PTX_2 UniProtKB/Swiss-Prot
RatGTEx ENSRNOG00000016156 RatGTEx
SMART PTX UniProtKB/Swiss-Prot
Superfamily-SCOP SSF49899 UniProtKB/Swiss-Prot
UniProt A6HSU2_RAT UniProtKB/TrEMBL
  F1LSY2_RAT UniProtKB/TrEMBL
  NPTXR_RAT UniProtKB/Swiss-Prot, ENTREZGENE


Nomenclature History
Date Current Symbol Current Name Previous Symbol Previous Name Description Reference Status
2005-01-20 Nptxr  neuronal pentraxin receptor      Symbol and Name status set to approved 1299863 APPROVED
2003-02-27 Nptxr  neuronal pentraxin receptor      Symbol and Name status set to provisional 70820 PROVISIONAL

RGD Curation Notes
Note Type Note Reference
gene_protein putative integral membrane pentraxin 633357