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Gene: Gmpr2 (guanosine monophosphate reductase 2) Rattus norvegicus

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Symbol:

Gmpr2

Name:

guanosine monophosphate reductase 2

RGD ID:

628875

Description:

Predicted to enable GMP reductase activity. Predicted to be involved in GMP metabolic process. Predicted to be part of GMP reductase complex. Orthologous to human GMPR2 (guanosine monophosphate reductase 2); INTERACTS WITH 1,3-dinitrobenzene; 2,3,7,8-tetrachlorodibenzodioxine; 2,4-dinitrotoluene.

Type:

protein-coding

RefSeq Status:

VALIDATED

Previously known as:

GMP reductase 2; guanosine monophosphate reductase isolog; MGC108877

RGD Orthologs

Alliance Orthologs

More Info

more info ...

More Info

Species	Gene symbol and name	Data Source	Assertion derived from	less info ...
Orthologs ¹				less info ...
Homo sapiens (human):	GMPR2 (guanosine monophosphate reductase 2)	HGNC	Ensembl, HomoloGene, NCBI, OMA, OrthoDB, Panther, PhylomeDB
Mus musculus (house mouse):	Gmpr2 (guanosine monophosphate reductase 2)	Transitive Ortholog Pipeline	Transitive Ortholog Pipeline
Chinchilla lanigera (long-tailed chinchilla):	Gmpr2 (guanosine monophosphate reductase 2)	Transitive Ortholog Pipeline	Transitive Ortholog Pipeline
Pan paniscus (bonobo/pygmy chimpanzee):	GMPR2 (guanosine monophosphate reductase 2)	Transitive Ortholog Pipeline	Transitive Ortholog Pipeline
Canis lupus familiaris (dog):	GMPR2 (guanosine monophosphate reductase 2)	Transitive Ortholog Pipeline	Transitive Ortholog Pipeline
Ictidomys tridecemlineatus (thirteen-lined ground squirrel):	Gmpr2 (guanosine monophosphate reductase 2)	Transitive Ortholog Pipeline	Transitive Ortholog Pipeline
Sus scrofa (pig):	GMPR2 (guanosine monophosphate reductase 2)	Transitive Ortholog Pipeline	Transitive Ortholog Pipeline
Chlorocebus sabaeus (green monkey):	GMPR2 (guanosine monophosphate reductase 2)	Transitive Ortholog Pipeline	Transitive Ortholog Pipeline
Heterocephalus glaber (naked mole-rat):	Gmpr2 (guanosine monophosphate reductase 2)	Transitive Ortholog Pipeline	Transitive Ortholog Pipeline
Rattus rattus (black rat):	Gmpr2 (guanosine monophosphate reductase 2)	Transitive Ortholog Pipeline	Transitive Ortholog Pipeline
Other homologs ²
Homo sapiens (human):	GMPR (guanosine monophosphate reductase)	HGNC	OrthoDB
Alliance orthologs ³
Homo sapiens (human):	GMPR2 (guanosine monophosphate reductase 2)	Alliance	DIOPT (Ensembl Compara\|HGNC\|Hieranoid\|InParanoid\|OMA\|OrthoFinder\|OrthoInspector\|PANTHER\|PhylomeDB\|SonicParanoid)
Homo sapiens (human):	GMPR2 (guanosine monophosphate reductase 2)	Alliance	DIOPT (Ensembl Compara\|HGNC\|Hieranoid\|OMA\|OrthoFinder\|PANTHER\|PhylomeDB)
Mus musculus (house mouse):	Gmpr2 (guanosine monophosphate reductase 2)	Alliance	DIOPT (Ensembl Compara\|Hieranoid\|InParanoid\|OMA\|OrthoFinder\|OrthoInspector\|PANTHER\|PhylomeDB\|SonicParanoid)
Mus musculus (house mouse):	Gmpr2 (guanosine monophosphate reductase 2)	Alliance	DIOPT (Ensembl Compara\|Hieranoid\|InParanoid\|OMA\|OrthoFinder\|PANTHER\|PhylomeDB\|SonicParanoid)
Danio rerio (zebrafish):	gmpr2 (guanosine monophosphate reductase 2)	Alliance	DIOPT (Ensembl Compara\|Hieranoid\|InParanoid\|OMA\|OrthoFinder\|PANTHER\|SonicParanoid)
Danio rerio (zebrafish):	gmpr2 (guanosine monophosphate reductase 2)	Alliance	DIOPT (Ensembl Compara\|Hieranoid\|OMA\|OrthoFinder\|PANTHER\|PhylomeDB)
Caenorhabditis elegans (roundworm):	F32D1.5	Alliance	DIOPT (Hieranoid\|InParanoid\|OMA\|OrthoFinder\|OrthoInspector\|PANTHER\|PhylomeDB\|SonicParanoid)
Caenorhabditis elegans (roundworm):	F32D1.5	Alliance	DIOPT (Hieranoid\|OMA\|OrthoFinder\|PANTHER\|PhylomeDB)
Xenopus tropicalis (tropical clawed frog):	gmpr2	Alliance	DIOPT (Ensembl Compara\|Hieranoid\|InParanoid\|OMA\|OrthoFinder\|OrthoInspector\|PANTHER\|SonicParanoid)
Xenopus tropicalis (tropical clawed frog):	gmpr2	Alliance	DIOPT (Ensembl Compara\|Hieranoid\|OMA\|OrthoFinder\|PANTHER\|PhylomeDB)

Latest Assembly:

GRCr8 - GRCr8 Assembly

Position:

Rat Assembly	Chr	Position (strand)	Source	Genome Browsers
Rat Assembly	Chr	Position (strand)	Source	JBrowse	NCBI	UCSC	Ensembl
GRCr8	15	33,135,455 - 33,144,923 (+)	NCBI	GRCr8	GRCr8		GRCr8
GRCr8 Ensembl	15	33,135,682 - 33,144,899 (+)	Ensembl				GRCr8
mRatBN7.2	15	29,165,421 - 29,175,933 (+)	NCBI	mRatBN7.2	mRatBN7.2
mRatBN7.2 Ensembl	15	29,165,783 - 29,174,935 (+)	Ensembl				mRatBN7.2
UTH_Rnor_SHR_Utx	15	31,011,501 - 31,020,661 (+)	NCBI	UTH_Rnor_SHR_Utx	UTH_Rnor_SHR_Utx
UTH_Rnor_SHRSP_BbbUtx_1.0	15	32,158,722 - 32,167,882 (+)	NCBI	UTH_Rnor_SHRSP_BbbUtx_1.0	UTH_Rnor_SHRSP_BbbUtx_1.0
UTH_Rnor_WKY_Bbb_1.0	15	30,401,065 - 30,410,221 (+)	NCBI	UTH_Rnor_WKY_Bbb_1.0	UTH_Rnor_WKY_Bbb_1.0
Rnor_6.0	15	34,352,857 - 34,362,074 (+)	NCBI	Rnor_6.0	Rnor_6.0	rn6	Rnor6.0
Rnor_6.0 Ensembl	15	34,352,914 - 34,362,032 (+)	Ensembl			rn6	Rnor6.0
Rnor_5.0	15	38,242,834 - 38,252,051 (+)	NCBI	Rnor_5.0	Rnor_5.0	rn5
RGSC_v3.4	15	33,820,865 - 33,830,025 (+)	NCBI	RGSC_v3.4	RGSC_v3.4	rn4
Celera	15	28,742,209 - 28,751,367 (+)	NCBI		Celera
RGSC_v3.1	15	33,836,608 - 33,840,631 (+)	NCBI
Cytogenetic Map	15	p13	NCBI

JBrowse:

View Region in Genome Browser (JBrowse)

Annotation Click to see Annotation Detail View

Disease Annotations Click to see Annotation Detail View

adenocarcinoma (ISO)

Stomach Neoplasms (ISO)

Gene-Chemical Interaction Annotations Click to see Annotation Detail View

1,2-dimethylhydrazine (ISO)

1,3-dinitrobenzene (EXP)

2,2',4,4'-Tetrabromodiphenyl ether (ISO)

2,3,7,8-tetrachlorodibenzodioxine (EXP,ISO)

2,4,6-tribromophenol (ISO)

2,4-dinitrotoluene (EXP)

2,6-dinitrotoluene (EXP)

2-methylcholine (ISO)

3,3',5,5'-tetrabromobisphenol A (ISO)

3-isobutyl-1-methyl-7H-xanthine (ISO)

4,4'-diaminodiphenylmethane (ISO)

acrolein (ISO)

acrylamide (EXP)

aflatoxin B1 (ISO)

alpha-pinene (ISO)

ammonium chloride (EXP)

Aroclor 1254 (ISO)

arsenite(3-) (ISO)

benzo[a]pyrene (ISO)

bisphenol A (EXP,ISO)

bisphenol AF (ISO)

cadmium atom (ISO)

cadmium dichloride (ISO)

carbamazepine (ISO)

carbon nanotube (ISO)

cisplatin (ISO)

cobalt dichloride (EXP)

copper(II) sulfate (ISO)

cyclosporin A (ISO)

deguelin (ISO)

dexamethasone (ISO)

Dibutyl phosphate (ISO)

dicrotophos (ISO)

dioxygen (ISO)

doxorubicin (ISO)

elemental selenium (ISO)

fipronil (EXP)

flutamide (EXP)

folic acid (ISO)

FR900359 (ISO)

indometacin (ISO)

ivermectin (ISO)

leflunomide (EXP)

methylmercury chloride (ISO)

nimesulide (EXP)

okadaic acid (ISO)

ozone (ISO)

paracetamol (ISO)

pentachlorophenol (ISO)

rotenone (ISO)

selenium atom (ISO)

Soman (EXP)

sulforaphane (ISO)

tert-butyl hydroperoxide (ISO)

tetracycline (ISO)

thiram (ISO)

triphenyl phosphate (ISO)

Gene Ontology Annotations Click to see Annotation Detail View

Biological Process

GMP metabolic process (IEA,ISO)

nucleotide metabolic process (IEA)

purine nucleobase metabolic process (IEA)

purine nucleotide metabolic process (IEA)

Cellular Component

GMP reductase complex (IEA)

Molecular Function

catalytic activity (IEA)

GMP reductase activity (IEA,ISO)

metal ion binding (IEA)

oxidoreductase activity (IEA)

protein binding (ISO)

References

References - curated

#	Reference Title	Reference Citation
1.	Phylogenetic-based propagation of functional annotations within the Gene Ontology consortium.	Gaudet P, etal., Brief Bioinform. 2011 Sep;12(5):449-62. doi: 10.1093/bib/bbr042. Epub 2011 Aug 27.
2.	Electronic Transfer of LocusLink and RefSeq Data	NCBI rat LocusLink and RefSeq merged data July 26, 2002
3.	GOA pipeline	RGD automated data pipeline
4.	ClinVar Automated Import and Annotation Pipeline	RGD automated import pipeline for ClinVar variants, variant-to-disease annotations and gene-to-disease annotations
5.	Data Import for Chemical-Gene Interactions	RGD automated import pipeline for gene-chemical interactions
6.	Cloning and functional characterization of GMPR2, a novel human guanosine monophosphate reductase, which promotes the monocytic differentiation of HL-60 leukemia cells.	Zhang J, etal., J Cancer Res Clin Oncol 2003 Feb;129(2):76-83. Epub 2003 Mar 1.

Additional References at PubMed

PMID:218932	PMID:12009299	PMID:12477932	PMID:16778019	PMID:22037469

Genomics

Comparative Map Data

Gmpr2
(Rattus norvegicus - Norway rat)

Rat Assembly	Chr	Position (strand)	Source	Genome Browsers
Rat Assembly	Chr	Position (strand)	Source	JBrowse	NCBI	UCSC	Ensembl
GRCr8	15	33,135,455 - 33,144,923 (+)	NCBI	GRCr8	GRCr8		GRCr8
GRCr8 Ensembl	15	33,135,682 - 33,144,899 (+)	Ensembl				GRCr8
mRatBN7.2	15	29,165,421 - 29,175,933 (+)	NCBI	mRatBN7.2	mRatBN7.2
mRatBN7.2 Ensembl	15	29,165,783 - 29,174,935 (+)	Ensembl				mRatBN7.2
UTH_Rnor_SHR_Utx	15	31,011,501 - 31,020,661 (+)	NCBI	UTH_Rnor_SHR_Utx	UTH_Rnor_SHR_Utx
UTH_Rnor_SHRSP_BbbUtx_1.0	15	32,158,722 - 32,167,882 (+)	NCBI	UTH_Rnor_SHRSP_BbbUtx_1.0	UTH_Rnor_SHRSP_BbbUtx_1.0
UTH_Rnor_WKY_Bbb_1.0	15	30,401,065 - 30,410,221 (+)	NCBI	UTH_Rnor_WKY_Bbb_1.0	UTH_Rnor_WKY_Bbb_1.0
Rnor_6.0	15	34,352,857 - 34,362,074 (+)	NCBI	Rnor_6.0	Rnor_6.0	rn6	Rnor6.0
Rnor_6.0 Ensembl	15	34,352,914 - 34,362,032 (+)	Ensembl			rn6	Rnor6.0
Rnor_5.0	15	38,242,834 - 38,252,051 (+)	NCBI	Rnor_5.0	Rnor_5.0	rn5
RGSC_v3.4	15	33,820,865 - 33,830,025 (+)	NCBI	RGSC_v3.4	RGSC_v3.4	rn4
Celera	15	28,742,209 - 28,751,367 (+)	NCBI		Celera
RGSC_v3.1	15	33,836,608 - 33,840,631 (+)	NCBI
Cytogenetic Map	15	p13	NCBI

GMPR2
(Homo sapiens - human)

Human Assembly	Chr	Position (strand)	Source	Genome Browsers
Human Assembly	Chr	Position (strand)	Source	JBrowse	NCBI	UCSC	Ensembl
GRCh38	14	24,232,622 - 24,239,242 (+)	NCBI	GRCh38	GRCh38	hg38	GRCh38
GRCh38.p14 Ensembl	14	24,232,211 - 24,239,246 (+)	Ensembl			hg38	GRCh38
GRCh37	14	24,701,828 - 24,708,448 (+)	NCBI	GRCh37	GRCh37	hg19	GRCh37
Build 36	14	23,771,488 - 23,778,288 (+)	NCBI	Build 36	Build 36	hg18	NCBI36
Build 34	14	23,771,942 - 23,778,285	NCBI
Celera	14	4,566,286 - 4,573,086 (+)	NCBI		Celera
Cytogenetic Map	14	q12	NCBI
HuRef	14	4,816,418 - 4,823,218 (+)	NCBI		HuRef
CHM1_1	14	24,700,271 - 24,707,071 (+)	NCBI		CHM1_1
T2T-CHM13v2.0	14	18,430,995 - 18,437,615 (+)	NCBI		T2T-CHM13v2.0

Gmpr2
(Mus musculus - house mouse)

Mouse Assembly	Chr	Position (strand)	Source	Genome Browsers
Mouse Assembly	Chr	Position (strand)	Source	JBrowse	NCBI	UCSC	Ensembl
GRCm39	14	55,909,692 - 55,935,602 (+)	NCBI	GRCm39	GRCm39	mm39
GRCm39 Ensembl	14	55,909,398 - 55,916,657 (+)	Ensembl		GRCm39 Ensembl		GRCm39
GRCm38	14	55,672,235 - 55,687,861 (+)	NCBI	GRCm38	GRCm38	mm10	GRCm38
GRCm38.p6 Ensembl	14	55,671,941 - 55,679,200 (+)	Ensembl			mm10	GRCm38
MGSCv37	14	56,291,072 - 56,297,588 (+)	NCBI	MGSCv37	MGSCv37	mm9	NCBIm37
MGSCv36	14	54,626,307 - 54,633,271 (+)	NCBI		MGSCv36	mm8
Celera	14	53,477,308 - 53,483,824 (+)	NCBI		Celera
Cytogenetic Map	14	C3	NCBI
cM Map	14	28.19	NCBI

Gmpr2
(Chinchilla lanigera - long-tailed chinchilla)

Chinchilla Assembly	Chr	Position (strand)	Source	Genome Browsers
Chinchilla Assembly	Chr	Position (strand)	Source	JBrowse	NCBI	UCSC	Ensembl
ChiLan1.0 Ensembl	NW_004955409	36,166,129 - 36,173,245 (-)	Ensembl
ChiLan1.0	NW_004955409	36,166,129 - 36,173,247 (-)	NCBI	ChiLan1.0	ChiLan1.0

GMPR2
(Pan paniscus - bonobo/pygmy chimpanzee)

Bonobo Assembly	Chr	Position (strand)	Source	Genome Browsers
Bonobo Assembly	Chr	Position (strand)	Source	JBrowse	NCBI	UCSC	Ensembl
NHGRI_mPanPan1-v2	15	25,586,650 - 25,594,233 (+)	NCBI		NHGRI_mPanPan1-v2
NHGRI_mPanPan1	14	24,803,398 - 24,810,725 (+)	NCBI		NHGRI_mPanPan1
Mhudiblu_PPA_v0	14	5,008,664 - 5,015,461 (+)	NCBI	Mhudiblu_PPA_v0	Mhudiblu_PPA_v0	panPan3
PanPan1.1	14	23,132,227 - 23,139,044 (+)	NCBI	PanPan1.1	PanPan1.1	panPan2
PanPan1.1 Ensembl	14	23,132,227 - 23,139,044 (+)	Ensembl			panPan2	panpan1.1

GMPR2
(Canis lupus familiaris - dog)

Dog Assembly	Chr	Position (strand)	Source	Genome Browsers
Dog Assembly	Chr	Position (strand)	Source	JBrowse	NCBI	UCSC	Ensembl
CanFam3.1	8	4,213,238 - 4,220,740 (+)	NCBI	CanFam3.1	CanFam3.1	canFam3	CanFam3.1
CanFam3.1 Ensembl	8	4,213,373 - 4,220,740 (+)	Ensembl			canFam3	CanFam3.1
Dog10K_Boxer_Tasha	8	4,134,799 - 4,142,420 (+)	NCBI	Dog10K_Boxer_Tasha	Dog10K_Boxer_Tasha
ROS_Cfam_1.0	8	4,323,889 - 4,331,513 (+)	NCBI	ROS_Cfam_1.0	ROS_Cfam_1.0
ROS_Cfam_1.0 Ensembl	8	4,324,117 - 4,331,509 (+)	Ensembl				ROS_Cfam_1.0 Ensembl
UMICH_Zoey_3.1	8	4,014,274 - 4,021,895 (+)	NCBI		UMICH_Zoey_3.1
UNSW_CanFamBas_1.0	8	4,076,063 - 4,083,696 (+)	NCBI		UNSW_CanFamBas_1.0
UU_Cfam_GSD_1.0	8	4,338,401 - 4,346,026 (+)	NCBI	UU_Cfam_GSD_1.0	UU_Cfam_GSD_1.0

Gmpr2
(Ictidomys tridecemlineatus - thirteen-lined ground squirrel)

Squirrel Assembly	Chr	Position (strand)	Source	Genome Browsers
Squirrel Assembly	Chr	Position (strand)	Source	JBrowse	NCBI	UCSC	Ensembl
HiC_Itri_2	NW_024408640	36,058,339 - 36,065,069 (+)	NCBI		HiC_Itri_2
SpeTri2.0 Ensembl	NW_004936722	324,301 - 330,309 (-)	Ensembl		SpeTri2.0 Ensembl
SpeTri2.0	NW_004936722	323,917 - 330,648 (-)	NCBI	SpeTri2.0	SpeTri2.0		SpeTri2.0

GMPR2
(Sus scrofa - pig)

Pig Assembly	Chr	Position (strand)	Source	Genome Browsers
Pig Assembly	Chr	Position (strand)	Source	JBrowse	NCBI	UCSC	Ensembl
Sscrofa11.1 Ensembl	7	75,060,676 - 75,070,084 (-)	Ensembl			susScr11	Sscrofa11.1
Sscrofa11.1	7	75,060,680 - 75,070,079 (-)	NCBI	Sscrofa11.1	Sscrofa11.1	susScr11	Sscrofa11.1
Sscrofa10.2	7	80,331,603 - 80,340,768 (-)	NCBI	Sscrofa10.2	Sscrofa10.2	susScr3

GMPR2
(Chlorocebus sabaeus - green monkey)

Green Monkey Assembly	Chr	Position (strand)	Source	Genome Browsers
Green Monkey Assembly	Chr	Position (strand)	Source	JBrowse	NCBI	UCSC	Ensembl
ChlSab1.1	24	1,215,137 - 1,222,219 (+)	NCBI	ChlSab1.1	ChlSab1.1	chlSab2
ChlSab1.1 Ensembl	24	1,215,357 - 1,225,716 (+)	Ensembl		ChlSab1.1 Ensembl	chlSab2	Vervet-AGM
Vero_WHO_p1.0	NW_023666036	12,899,552 - 12,906,635 (+)	NCBI	Vero_WHO_p1.0	Vero_WHO_p1.0

Gmpr2
(Heterocephalus glaber - naked mole-rat)

Naked Mole-Rat Assembly	Chr	Position (strand)	Source	Genome Browsers
Naked Mole-Rat Assembly	Chr	Position (strand)	Source	JBrowse	NCBI	UCSC	Ensembl
HetGla_female_1.0 Ensembl	NW_004624820	8,312,386 - 8,319,799 (-)	Ensembl		HetGla_female_1.0 Ensembl	hetGla2	HetGla_female_1.0 Ensembl
HetGla 1.0	NW_004624820	8,312,253 - 8,319,585 (-)	NCBI	HetGla 1.0	HetGla 1.0	hetGla2

Gmpr2
(Rattus rattus - black rat)

Black Rat Assembly	Chr	Position (strand)	Source	Genome Browsers
Black Rat Assembly	Chr	Position (strand)	Source	JBrowse	NCBI	UCSC	Ensembl
Rrattus_CSIRO_v1	12	69,326,153 - 69,332,886 (-)	NCBI		Rrattus_CSIRO_v1

Variants

Variants in Gmpr2
23 total Variants

miRNA Target Status (No longer updated)

Predicted Target Of

	Summary Value
Count of predictions:	544
Count of miRNA genes:	210
Interacting mature miRNAs:	240
Transcripts:	ENSRNOT00000067150
Prediction methods:	Microtar, Miranda, Rnahybrid
Result types:	miRGate_prediction

The detailed report is available here: Full Report CSV TAB Printer

miRNA Target Status data imported from miRGate (http://mirgate.bioinfo.cnio.es/).
For more information about miRGate, see PMID:25858286 or access the full paper here.

QTLs in Region (GRCr8)

The following QTLs overlap with this region.

RGD ID	Symbol	Name	LOD	P Value	Trait	Sub Trait	Chr	Start	Stop	Species
1641913	Colcr2	Colorectal carcinoma resistance QTL 2	6.57	0.0197	intestine integrity trait (VT:0010554)	poorly differentiated malignant colorectal tumor number (CMO:0002076)	15	2315759	47315759	Rat
5684946	Bss98	Bone structure and strength QTL 98	3.9	0.0026	tibia strength trait (VT:1000284)	tibia ultimate force (CMO:0001734)	15	1	39411584	Rat
1641887	Alcrsp14	Alcohol response QTL 14			response to alcohol trait (VT:0010489)	brain neurotensin receptor 1 density (CMO:0002068)	15	1	44836456	Rat
738017	Hcas7	Hepatocarcinoma susceptibility QTL 7	2.91		liver integrity trait (VT:0010547)	liver nonremodeling tumorous lesion volume to total liver volume ratio (CMO:0001464)	15	2315759	53331089	Rat
152025253	Hrtrt24	Heart rate QTL 24	3.82		heart pumping trait (VT:2000009)		15	31855803	92671517	Rat
10755503	Bp391	Blood pressure QTL 391	2.37		arterial blood pressure trait (VT:2000000)	systolic blood pressure (CMO:0000004)	15	22728353	42072050	Rat
10054130	Srcrt8	Stress Responsive Cort QTL 8	2.18	0.0085	blood corticosterone amount (VT:0005345)	plasma corticosterone level (CMO:0001173)	15	28527732	73527732	Rat
1578646	Bmd18	Bone mineral density QTL 18	5.2		femur mineral mass (VT:0010011)	trabecular volumetric bone mineral density (CMO:0001729)	15	25285835	104695021	Rat
2324620	Coatc3	Coat color QTL 3			coat/hair pigmentation trait (VT:0010463)	pigmented coat/hair area to total coat/hair area ratio (CMO:0001810)	15	22336351	52597086	Rat
1578647	Bmd17	Bone mineral density QTL 17	4		femur mineral mass (VT:0010011)	total volumetric bone mineral density (CMO:0001728)	15	25285835	104695021	Rat
61424	Scl1	Serum cholesterol level QTL 1	7.7	0.001	blood cholesterol amount (VT:0000180)	serum total cholesterol level (CMO:0000363)	15	19155762	87086765	Rat
10401805	Kidm51	Kidney mass QTL 51			kidney mass (VT:0002707)	both kidneys wet weight (CMO:0000085)	15	2785923	47785923	Rat
2317750	Glom26	Glomerulus QTL 26	4.3		urine protein amount (VT:0005160)	urine protein level (CMO:0000591)	15	14926594	71614418	Rat
631550	Bw7	Body weight QTL 7	3.6		body mass (VT:0001259)	body weight (CMO:0000012)	15	22336351	67336351	Rat
1578660	Bss19	Bone structure and strength QTL 19	4.3		femur morphology trait (VT:0000559)	bone trabecular cross-sectional area (CMO:0002311)	15	25285835	104695021	Rat

Markers in Region

RH128726

Rat Assembly	Chr	Position (strand)	Source	JBrowse
mRatBN7.2	15	29,173,250 - 29,173,445 (+)	MAPPER	mRatBN7.2
Rnor_6.0	15	34,360,348 - 34,360,542	NCBI	Rnor6.0
Rnor_5.0	15	38,250,325 - 38,250,519	UniSTS	Rnor5.0
RGSC_v3.4	15	33,828,299 - 33,828,493	UniSTS	RGSC3.4
Celera	15	28,749,641 - 28,749,835	UniSTS
RH 3.4 Map	15	260.81	UniSTS
Cytogenetic Map	15	p13	UniSTS

RH129602

Rat Assembly	Chr	Position (strand)	Source	JBrowse
GRCr8	15	33,143,954 - 33,144,570 (+)	Marker Load Pipeline
mRatBN7.2	15	29,174,008 - 29,174,624 (+)	MAPPER	mRatBN7.2
Rnor_6.0	15	34,361,106 - 34,361,721	NCBI	Rnor6.0
Rnor_5.0	15	38,251,083 - 38,251,698	UniSTS	Rnor5.0
RGSC_v3.4	15	33,829,057 - 33,829,672	UniSTS	RGSC3.4
Celera	15	28,750,399 - 28,751,014	UniSTS
RH 3.4 Map	15	213.7	UniSTS
Cytogenetic Map	15	p13	UniSTS

RH129694

Rat Assembly	Chr	Position (strand)	Source	JBrowse
mRatBN7.2	15	29,173,871 - 29,174,054 (+)	MAPPER	mRatBN7.2
Rnor_6.0	15	34,360,969 - 34,361,151	NCBI	Rnor6.0
Rnor_5.0	15	38,250,946 - 38,251,128	UniSTS	Rnor5.0
RGSC_v3.4	15	33,828,920 - 33,829,102	UniSTS	RGSC3.4
Celera	15	28,750,262 - 28,750,444	UniSTS
RH 3.4 Map	15	213.69	UniSTS
Cytogenetic Map	15	p13	UniSTS

AW530559

Rat Assembly	Chr	Position (strand)	Source	JBrowse
mRatBN7.2	15	29,170,742 - 29,170,902 (+)	MAPPER	mRatBN7.2
Rnor_6.0	15	34,357,840 - 34,357,999	NCBI	Rnor6.0
Rnor_5.0	15	38,247,817 - 38,247,976	UniSTS	Rnor5.0
RGSC_v3.4	15	33,825,791 - 33,825,950	UniSTS	RGSC3.4
Celera	15	28,747,133 - 28,747,292	UniSTS
RH 3.4 Map	15	261.01	UniSTS
Cytogenetic Map	15	p13	UniSTS

Expression

RNA-SEQ Expression

Click on a value in the shaded box below the category label to view a detailed expression data table for that system.

Download All Expressed Objects for this Gene

alimentary part of gastrointestinal system	appendage	circulatory system	ectoderm	endocrine system	endoderm	exocrine system	hemolymphoid system	hepatobiliary system	integumental system	mesenchyme	mesoderm	musculoskeletal system	nervous system	renal system	reproductive system	respiratory system	sensory system	visual system
16	12	67	165	91	90	59	92	59	6	356	192	11	144	81	92	31	17	17

Sequence

Nucleotide Sequences


RefSeq Transcripts	NM_001013036	(Get FASTA)	NCBI Sequence Viewer	Search GEO for Microarray Profiles
	NM_001413771	(Get FASTA)	NCBI Sequence Viewer	Search GEO for Microarray Profiles
	NM_001413772	(Get FASTA)	NCBI Sequence Viewer	Search GEO for Microarray Profiles
	XM_006251926	(Get FASTA)	NCBI Sequence Viewer	Search GEO for Microarray Profiles
	XM_063273959	(Get FASTA)	NCBI Sequence Viewer	Search GEO for Microarray Profiles
	XM_063273960	(Get FASTA)	NCBI Sequence Viewer	Search GEO for Microarray Profiles
	XM_063273961	(Get FASTA)	NCBI Sequence Viewer	Search GEO for Microarray Profiles
	XM_063273963	(Get FASTA)	NCBI Sequence Viewer	Search GEO for Microarray Profiles
GenBank Nucleotide	AC116083	(Get FASTA)	NCBI Sequence Viewer	Search GEO for Microarray Profiles
	AF350372	(Get FASTA)	NCBI Sequence Viewer	Search GEO for Microarray Profiles
	BC089848	(Get FASTA)	NCBI Sequence Viewer	Search GEO for Microarray Profiles
	CH474049	(Get FASTA)	NCBI Sequence Viewer	Search GEO for Microarray Profiles
	DQ758870	(Get FASTA)	NCBI Sequence Viewer	Search GEO for Microarray Profiles
	FQ211507	(Get FASTA)	NCBI Sequence Viewer	Search GEO for Microarray Profiles
	JAXUCZ010000015	(Get FASTA)	NCBI Sequence Viewer	Search GEO for Microarray Profiles

Ensembl Acc Id:

ENSRNOT00000067150 ⟹ ENSRNOP00000060006

Type:

CODING

Position:

Rat Assembly	Chr	Position (strand)	Source
GRCr8 Ensembl	15	33,135,745 - 33,144,884 (+)	Ensembl
mRatBN7.2 Ensembl	15	29,165,816 - 29,174,935 (+)	Ensembl
Rnor_6.0 Ensembl	15	34,352,914 - 34,362,032 (+)	Ensembl

Ensembl Acc Id:

ENSRNOT00000084554 ⟹ ENSRNOP00000070108

Type:

CODING

Position:

Rat Assembly	Chr	Position (strand)	Source
GRCr8 Ensembl	15	33,135,682 - 33,144,899 (+)	Ensembl
mRatBN7.2 Ensembl	15	29,165,783 - 29,173,635 (+)	Ensembl
Rnor_6.0 Ensembl	15	34,353,495 - 34,360,417 (+)	Ensembl

RefSeq Acc Id:

NM_001013036 ⟹ NP_001013054

RefSeq Status:

VALIDATED

Type:

CODING

Position:

Rat Assembly	Chr	Position (strand)	Source
GRCr8	15	33,135,763 - 33,144,923 (+)	NCBI
mRatBN7.2	15	29,165,816 - 29,174,977 (+)	NCBI
Rnor_6.0	15	34,352,914 - 34,362,074 (+)	NCBI
Rnor_5.0	15	38,242,834 - 38,252,051 (+)	NCBI
RGSC_v3.4	15	33,820,865 - 33,830,025 (+)	RGD
Celera	15	28,742,209 - 28,751,367 (+)	RGD

Sequence:

show sequence

CGAGGCCCCACGCGGCTAGTTGCTAAGCAAAAGAATCTTTGGTCAACTGCCATCTGAGTCAAAG
CAACAGGAACACTTCATCATGCCTCATATTGACAATGATGTGAAACTAGACTTCAAGGATGTCC
TGTTGAGGCCCAAACGCAGCACTCTAAAGTCTCGAAGTGAGGTAGATCTCACAAGATCATTTTC
ATTCCGGAACTCAAAGCAGATGTACAGTGGCATCCCCATCATTGCTGCCAATATGGATACCGTG
GGTACTTTTGAAATGGCTAAGGTTCTCTGTAAGTTTTCCCTGTTCACTGCCATCCATAAGCACT
ACAGCATCCATCAGTGGCAAGAGTTTGCTAACCAGAATCCTGACTGTCTTGAGTATCTAGCTGC
CAGCTCAGGCACAGGCTCTGCTGACTTTGAGCAGCTGGAACAGATCCTAGAAGCTATTCCCCAA
GTCAAATATATATGCCTGGATGTGGCTAATGGCTACTCAGAACATTTTGTTGAATCCGTAAAGG
ATGTACGGAAGCGATTCCCCCAACACACCATCATGGCAGGCAATGTGGTAACAGGAGAGATGGT
AGAAGAGCTAATCCTCTCTGGGGCTGACATCATCAAAGTGGGAATTGGTCCTGGCTCTGTCTGT
ACAACTCGGAAGAAAACTGGAGTTGGGTATCCACAGCTCAGTGCAGTGATGGAGTGTGCAGATG
CTGCTCATGGCCTCAAAGGCCACATCATTTCAGACGGAGGCTGCAGCTGCCCTGGAGATGTGGC
AAAGGCCTTTGGCGCAGGGGCTGACTTTGTGATGCTGGGTGGCATGCTGGCCGGGCACAGTGAA
TCAGGTGGTGAACTCATTGAGAGAAATGGCAAGAAATACAAGCTCTTCTATGGAATGAGTTCTG
AAATGGCTATGAAGAAGTATTCAGGAGGCGTGGCTGAGTACAGATAAATCTGAACCCTTTACAA
AGAAGTAAATATGAAAAGGGTCTCAACCAGCTCTAAGAGGAGCCTGGTATGTTTCTGAGGGATG
ACCCACACAGACTATGTAAGTATCAATTCCCCTTCTTCCCATCAAGACACTGAAAGAAATCAAG
ATTTAATTTTCAAGTGTTACAAAGCTTTTAGCAATGTAGCCAGGCAACTGAGGTCTCAGCTTCC
TCTTCTTTTCCACTAAACTTTTGCGTGCAGAGCTGCAGGGGTGGATGGACAGAGTGCAGAGGGT
ATTCCAGAGGACCTGAGTGAGAACTGTTTATGTAGCCAGAAAGGAATAGTCAGGACACTATGGA
ATGTAGTCACAAAACATGGGGATATAGGTGTCAAGCTTCGTCTTCTGATAGTTTTTCAAAGACT
CCTATAGAGAAGAAAGCATAATTCCTATAACTGGGAGCACTTCACACAGATTCCTCTTAACTGG
GAGCACTTCACACAGATTCCTCTAACTGGGATCACTTCACACAGATTCCTCTTAACTGGGAGCA
CTTCATACGGAGTCCTCTTAACTGGGAGCACTTCACACAGAGTCCTCTAACTGGGAGCACTTCA
CACAGATTCCTCTAACTGGGAGCACTTCACATAGATTCCTCTAACTGGGAGAACTTCACACAGA
TTCCTCTAACTGGGAGCACTTCACACGGAGTCCTCTTAACTGGGAGCACTTCACACAGATTCCT
CTAACTGGGAGAACTTCACACAGATTCCTCTAACTGGGAGCACTTCACACGGAGTCTTGCCTTA
TTCTGCCTATCCTCTGGCTCCACTTTCTGCTTCTTCCCACTCACCTTCCCTTCTTTCTGGTTAT
TTTCTTCCTCATCAGAGTCCAAAACCAAGTCCCCTATGGTAATAACAGAGCTACATAGGGACGG
GGATAGCACTGCAGGAAAGAAACCACAATTAAAACCACCTTTTTTTTTGGTTCTTTTTTTTCGG
AGCTGGGGACCGAACCCAGGGCCTTGCGCTTCCTAGGTAAGCGCTCTACCACTGAGCTAAATCC
CCAGCCCCTAAAACCACCTTTTTAGCCCAGCTTGGTAGCCCACACTTTTAATCCCTCCACTCTG
GAGGCTGAGGCAGGGGGATCTCTGAATTCAAGGCCAGCCTGGTCTATAGATAGAGTTCCTGGAC
AGCCAGGGTTATGAAGAAAAAAAATTATGTCAAAAAAAAAAAAAAAAAAAAAAAA

hide sequence

RefSeq Acc Id:

NM_001413771 ⟹ NP_001400700

RefSeq Status:

VALIDATED

Type:

CODING

Position:

Rat Assembly	Chr	Position (strand)	Source
GRCr8	15	33,135,739 - 33,143,431 (+)	NCBI
mRatBN7.2	15	29,165,792 - 29,173,485 (+)	NCBI

RefSeq Acc Id:

NM_001413772 ⟹ NP_001400701

RefSeq Status:

VALIDATED

Type:

CODING

Position:

Rat Assembly	Chr	Position (strand)	Source
GRCr8	15	33,135,739 - 33,143,431 (+)	NCBI
mRatBN7.2	15	29,165,792 - 29,173,485 (+)	NCBI

RefSeq Acc Id:

XM_006251926 ⟹ XP_006251988

Type:

CODING

Position:

Rat Assembly	Chr	Position (strand)	Source
GRCr8	15	33,135,696 - 33,143,425 (+)	NCBI
mRatBN7.2	15	29,165,421 - 29,173,479 (+)	NCBI
Rnor_6.0	15	34,352,857 - 34,360,579 (+)	NCBI
Rnor_5.0	15	38,242,834 - 38,252,051 (+)	NCBI

Sequence:

show sequence

CTTGGGTAGTCGGGTTTGCAATTAGCGGAATTTGTTCGAGTCGCCCTCAAGGCTCATCGAGGCC
CCACGCGGCTAGTTGCTAAGCAAAAGAATCTTTGGTCAACTGCCATCTGAGTCAAAGCAACAGG
AACACTTCATCATGCCTCATATTGACAATGATGTGAAACTAGACTTCAAGGATGTCCTGTTGAG
GCCCAAACGCAGCACTCTAAAGTCTCGAAGTGAGGTAGATCTCACAAGATCATTTTCATTCCGG
AACTCAAAGCAGATGTACAGTGGCATCCCCATCATTGCTGCCAATATGGATACCGTGGGTACTT
TTGAAATGGCTAAGGTTCTCTGTAAGTTTTCCCTGTTCACTGCCATCCATAAGCACTACAGCAT
CCATCAGTGGCAAGAGTTTGCTAACCAGAATCCTGACTGTCTTGAGTATCTAGCTGCCAGCTCA
GGCACAGGCTCTGCTGACTTTGAGCAGCTGGAACAGATCCTAGAAGCTATTCCCCAAGTCAAAT
ATATATGCCTGGATGTGGCTAATGGCTACTCAGAACATTTTGTTGAATCCGTAAAGGATGTACG
GAAGCGATTCCCCCAACACACCATCATGGCAGGCAATGTGGTAACAGGAGAGATGGTAGAAGAG
CTAATCCTCTCTGGGGCTGACATCATCAAAGTGGGAATTGGTCCTGGCTCTGTCTGTACAACTC
GGAAGAAAACTGGAGTTGGGTATCCACAGCTCAGTGCAGTGATGGAGTGTGCAGATGCTGCTCA
TGGCCTCAAAGGCCACATCATTTCAGACGGAGGCTGCAGCTGCCCTGGAGATGTGGCAAAGGCC
TTTGGCGCAGGGGCTGACTTTGTGATGCTGGGTGGCATGCTGGCCGGGCACAGTGAATCAGGTG
GTGAACTCATTGAGAGAAATGGCAAGAAATACAAGCTCTTCTATGGAATGAGTTCTGAAATGGC
TATGAAGAAGTATTCAGGAGGCGTGGCTGAGTACAGCTGCACATATAAAATGCTACGGGCACTA
TGGGAGAGGCAAGGAAGAAGACTTGAAATCAACAAATGAGAACCTGGGCACAGCCATGAAGATG
AGGGAAGAAAGAAGCTGATCCACACATAAATGGCAGACTTTGGTCTGGAAAAGGAAGGCTGTTG
GAATCATGTTAACCAGCTTCATTAAATAAGATCATCAAATGTCTAA

hide sequence

RefSeq Acc Id:

XM_063273959 ⟹ XP_063130029

Type:

CODING

Position:

Rat Assembly	Chr	Position (strand)	Source
GRCr8	15	33,135,824 - 33,143,425 (+)	NCBI

RefSeq Acc Id:

XM_063273960 ⟹ XP_063130030

Type:

CODING

Position:

Rat Assembly	Chr	Position (strand)	Source
GRCr8	15	33,135,455 - 33,143,425 (+)	NCBI

RefSeq Acc Id:

XM_063273961 ⟹ XP_063130031

Type:

CODING

Position:

Rat Assembly	Chr	Position (strand)	Source
GRCr8	15	33,135,824 - 33,143,425 (+)	NCBI

RefSeq Acc Id:

XM_063273963 ⟹ XP_063130033

Type:

CODING

Position:

Rat Assembly	Chr	Position (strand)	Source
GRCr8	15	33,136,343 - 33,143,425 (+)	NCBI

Protein Sequences


Protein RefSeqs	NP_001013054	(Get FASTA)	NCBI Sequence Viewer
	NP_001400700	(Get FASTA)	NCBI Sequence Viewer
	NP_001400701	(Get FASTA)	NCBI Sequence Viewer
	XP_006251988	(Get FASTA)	NCBI Sequence Viewer
	XP_063130029	(Get FASTA)	NCBI Sequence Viewer
	XP_063130030	(Get FASTA)	NCBI Sequence Viewer
	XP_063130031	(Get FASTA)	NCBI Sequence Viewer
	XP_063130033	(Get FASTA)	NCBI Sequence Viewer
GenBank Protein	AAH89848	(Get FASTA)	NCBI Sequence Viewer
	AAK31346	(Get FASTA)	NCBI Sequence Viewer
	EDM14265	(Get FASTA)	NCBI Sequence Viewer
Ensembl Protein	ENSRNOP00000060006
	ENSRNOP00000060006.2
	ENSRNOP00000070108
	ENSRNOP00000070108.3

RefSeq Acc Id:	NP_001013054 ⟸ NM_001013036
- Peptide Label:	isoform 1
- UniProtKB:	Q5FVP6 (UniProtKB/TrEMBL)
- Sequence:	show sequence MPHIDNDVKLDFKDVLLRPKRSTLKSRSEVDLTRSFSFRNSKQMYSGIPIIAANMDTVGTFEMA KVLCKFSLFTAIHKHYSIHQWQEFANQNPDCLEYLAASSGTGSADFEQLEQILEAIPQVKYICL DVANGYSEHFVESVKDVRKRFPQHTIMAGNVVTGEMVEELILSGADIIKVGIGPGSVCTTRKKT GVGYPQLSAVMECADAAHGLKGHIISDGGCSCPGDVAKAFGAGADFVMLGGMLAGHSESGGELI ERNGKKYKLFYGMSSEMAMKKYSGGVAEYR hide sequence

RefSeq Acc Id:

XP_006251988 ⟸ XM_006251926

- Peptide Label:

isoform X2

- Sequence:

show sequence

MPHIDNDVKLDFKDVLLRPKRSTLKSRSEVDLTRSFSFRNSKQMYSGIPIIAANMDTVGTFEMA
KVLCKFSLFTAIHKHYSIHQWQEFANQNPDCLEYLAASSGTGSADFEQLEQILEAIPQVKYICL
DVANGYSEHFVESVKDVRKRFPQHTIMAGNVVTGEMVEELILSGADIIKVGIGPGSVCTTRKKT
GVGYPQLSAVMECADAAHGLKGHIISDGGCSCPGDVAKAFGAGADFVMLGGMLAGHSESGGELI
ERNGKKYKLFYGMSSEMAMKKYSGGVAEYSCTYKMLRALWERQGRRLEINK

hide sequence

Ensembl Acc Id:

ENSRNOP00000060006 ⟸ ENSRNOT00000067150

Ensembl Acc Id:

ENSRNOP00000070108 ⟸ ENSRNOT00000084554

RefSeq Acc Id:	NP_001400701 ⟸ NM_001413772
- Peptide Label:	isoform 3

RefSeq Acc Id:	NP_001400700 ⟸ NM_001413771
- Peptide Label:	isoform 2
- UniProtKB:	A6KH36 (UniProtKB/TrEMBL)

RefSeq Acc Id:	XP_063130030 ⟸ XM_063273960
- Peptide Label:	isoform X1
- UniProtKB:	A6KH36 (UniProtKB/TrEMBL)

RefSeq Acc Id:	XP_063130029 ⟸ XM_063273959
- Peptide Label:	isoform X1
- UniProtKB:	A6KH36 (UniProtKB/TrEMBL)

RefSeq Acc Id:	XP_063130031 ⟸ XM_063273961
- Peptide Label:	isoform X2

RefSeq Acc Id:	XP_063130033 ⟸ XM_063273963
- Peptide Label:	isoform X3

Protein Domains

IMP dehydrogenase/GMP reductase

Protein Structures

Name	Modeler	Protein Id	AA Range	Protein Structure
AF-A0A0G2JX25-F1-model_v2	AlphaFold	A0A0G2JX25	1-348	view protein structure

Transcriptome

eQTL		View at Phenogen
WGCNA		View at Phenogen
Tissue/Strain Expression		View at Phenogen

Promoters

RGD ID:

13699683

Promoter ID:

EPDNEW_R10206

Type:

multiple initiation site

Name:

Gmpr2_1

Description:

guanosine monophosphate reductase 2

SO ACC ID:

SO:0000170

Source:

EPDNEW (Eukaryotic Promoter Database, http://epd.vital-it.ch/)

Experiment Methods:

Single-end sequencing.

Position:

Rat Assembly	Chr	Position (strand)	Source
Rnor_6.0	15	34,352,890 - 34,352,950	EPDNEW

Additional Information

External Database Links

Database	Acc Id	Source(s)
AGR Gene	RGD:628875	AgrOrtholog
BioCyc Gene	G2FUF-13698	BioCyc
Ensembl Genes	ENSRNOG00000020216	Ensembl, ENTREZGENE, UniProtKB/TrEMBL
Ensembl Transcript	ENSRNOT00000067150	ENTREZGENE
	ENSRNOT00000067150.5	UniProtKB/TrEMBL
	ENSRNOT00000084554	ENTREZGENE
	ENSRNOT00000084554.3	UniProtKB/TrEMBL
Gene3D-CATH	3.20.20.70	UniProtKB/TrEMBL
InterPro	Aldolase_TIM	UniProtKB/TrEMBL
	GMP_reduct1	UniProtKB/TrEMBL
	GMP_reductase	UniProtKB/TrEMBL
	IMP_DH/GMP_Rdtase_CS	UniProtKB/TrEMBL
	IMP_DH_GMPRt	UniProtKB/TrEMBL
KEGG Report	rno:192357	UniProtKB/TrEMBL
NCBI Gene	192357	ENTREZGENE
PANTHER	GMP REDUCTASE	UniProtKB/TrEMBL
	GMP REDUCTASE 2	UniProtKB/TrEMBL
Pfam	IMPDH	UniProtKB/TrEMBL
PhenoGen	Gmpr2	PhenoGen
PIRSF	GMP_reductase	UniProtKB/TrEMBL
PROSITE	IMP_DH_GMP_RED	UniProtKB/TrEMBL
RatGTEx	ENSRNOG00000020216	RatGTEx
SMART	IMPDH	UniProtKB/TrEMBL
Superfamily-SCOP	Inosine monophosphate dehydrogenase (IMPDH)	UniProtKB/TrEMBL
UniProt	A0A0G2JX25_RAT	UniProtKB/TrEMBL
	A6KH36	ENTREZGENE, UniProtKB/TrEMBL
	F7F3D1_RAT	UniProtKB/TrEMBL
	Q5FVP6	ENTREZGENE, UniProtKB/TrEMBL

Nomenclature History

Date	Current Symbol	Current Name	Previous Symbol	Previous Name	Description	Reference	Status
2003-04-09	Gmpr2	guanosine monophosphate reductase 2			Symbol and Name updated	629477	APPROVED
2003-03-14	Gmpr2	guanosine monophosphate reductase 2	Loc192357	guanosine monophosphate reductase isolog	Data merged from RGD:621848	628472	PROVISIONAL
2003-02-27	Gmpr2	guanosine monophosphate reductase 2			Symbol and Name status set to provisional	70820	PROVISIONAL
2002-08-07	Loc192357	guanosine monophosphate reductase isolog			Symbol and Name status set to provisional	70820	PROVISIONAL

Gene Annotator (Functional Annotation) unavailable	Gene Annotator (Annotation Distribution) unavailable	Variant Visualizer (Genomic Variants) unavailble Variant Visualizer (Genomic Variants) unavailable
Gene Annotator (Functional Annotation)	Gene Annotator (Annotation Distribution)	Variant Visualizer (Genomic Variants)

InterViewer (Protein-Protein Interactions) unavailable	Gviewer (Genome Viewer) unavailable	Variant Visualizer (Damaging Variants) unavailble Variant Visualizer (Damaging Variants) unavailable
InterViewer (Protein-Protein Interactions)	GViewer (Genome Viewer)	Variant Visualizer (Damaging Variants)

Gene Annotator (Annotation Comparison) unavailable	OLGA (Gene List Generator) unavailable
Gene Annotator (Annotation Comparison)	OLGA (Gene List Generator)	Excel (Download)

MOET (Multi-Ontology Enrichement) unavailable	GOLF (Gene-Ortholog Location Finder) unavailable
MOET (Multi-Ontology Enrichement)	GOLF (Gene-Ortholog Location Finder)

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Imported Disease Annotations - CTD