Atg101 (autophagy related 101) - Rat Genome Database

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Advanced Search (OLGA)
Gene: Atg101 (autophagy related 101) Rattus norvegicus
Analyze
Symbol: Atg101
Name: autophagy related 101
RGD ID: 1359310
Description: Predicted to enable identical protein binding activity and protein kinase binding activity. Predicted to be involved in autophagosome assembly; negative regulation of cell population proliferation; and positive regulation of autophagy. Predicted to be located in cytoplasm. Predicted to be part of Atg1/ULK1 kinase complex. Predicted to be active in phagophore assembly site. Orthologous to human ATG101 (autophagy related 101); INTERACTS WITH 2,3,7,8-tetrachlorodibenzodioxine; bisphenol A; endosulfan.
Type: protein-coding
RefSeq Status: VALIDATED
Previously known as: autophagy-related protein 101; MGC94201; RGD1359310; similar to RIKEN cDNA 9430023L20
RGD Orthologs
Human
Mouse
Chinchilla
Bonobo
Dog
Squirrel
Pig
Green Monkey
Naked Mole-Rat
Black Rat
Alliance Orthologs
More Info more info ...
Latest Assembly: GRCr8 - GRCr8 Assembly
Position:
Rat AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
GRCr87134,277,223 - 134,286,387 (+)NCBIGRCr8GRCr8GRCr8
GRCr8 Ensembl7134,272,847 - 134,286,379 (+)EnsemblGRCr8
mRatBN7.27132,398,477 - 132,407,643 (+)NCBImRatBN7.2mRatBN7.2
mRatBN7.2 Ensembl7132,398,483 - 132,407,633 (+)EnsemblmRatBN7.2
UTH_Rnor_SHR_Utx7134,200,027 - 134,209,527 (+)NCBIUTH_Rnor_SHR_UtxUTH_Rnor_SHR_Utx
UTH_Rnor_SHRSP_BbbUtx_1.07136,429,543 - 136,439,043 (+)NCBIUTH_Rnor_SHRSP_BbbUtx_1.0UTH_Rnor_SHRSP_BbbUtx_1.0
UTH_Rnor_WKY_Bbb_1.07136,339,980 - 136,348,905 (+)NCBIUTH_Rnor_WKY_Bbb_1.0UTH_Rnor_WKY_Bbb_1.0
Rnor_6.07142,929,381 - 142,938,559 (+)NCBIRnor_6.0Rnor_6.0rn6Rnor6.0
Rnor_6.0 Ensembl7142,929,400 - 142,938,554 (+)Ensemblrn6Rnor6.0
Rnor_5.07140,730,251 - 140,739,416 (+)NCBIRnor_5.0Rnor_5.0rn5
RGSC_v3.47140,029,813 - 140,038,925 (+)NCBIRGSC_v3.4RGSC_v3.4rn4
Celera7128,863,838 - 128,872,961 (+)NCBICelera
RGSC_v3.17140,106,249 - 140,115,362 (+)NCBI
Cytogenetic Map7q36NCBI
JBrowse:




Disease Annotations     Click to see Annotation Detail View

Gene Ontology Annotations     Click to see Annotation Detail View

Biological Process

Cellular Component

Molecular Function

References

References - curated
# Reference Title Reference Citation
1. Phylogenetic-based propagation of functional annotations within the Gene Ontology consortium. Gaudet P, etal., Brief Bioinform. 2011 Sep;12(5):449-62. doi: 10.1093/bib/bbr042. Epub 2011 Aug 27.
2. Data Import for Chemical-Gene Interactions RGD automated import pipeline for gene-chemical interactions
3. Generation and initial analysis of more than 15,000 full-length human and mouse cDNA sequences. Strausberg RL, etal., Proc Natl Acad Sci U S A. 2002 Dec 24;99(26):16899-903. Epub 2002 Dec 11.
4. Development and validation of prognostic model based on the analysis of autophagy-related genes in colon cancer. Wang Y, etal., Aging (Albany NY). 2021 Jul 27;13(14):19028-19047. doi: 10.18632/aging.203352. Epub 2021 Jul 27.
5. Comprehensive Analysis of the Potential Immune-Related Biomarker ATG101 that Regulates Apoptosis of Cholangiocarcinoma Cells After Photodynamic Therapy. Zhang ZJ, etal., Front Pharmacol. 2022 May 3;13:857774. doi: 10.3389/fphar.2022.857774. eCollection 2022.
Additional References at PubMed
PMID:19597335   PMID:20562859  


Genomics

Comparative Map Data
Atg101
(Rattus norvegicus - Norway rat)
Rat AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
GRCr87134,277,223 - 134,286,387 (+)NCBIGRCr8GRCr8GRCr8
GRCr8 Ensembl7134,272,847 - 134,286,379 (+)EnsemblGRCr8
mRatBN7.27132,398,477 - 132,407,643 (+)NCBImRatBN7.2mRatBN7.2
mRatBN7.2 Ensembl7132,398,483 - 132,407,633 (+)EnsemblmRatBN7.2
UTH_Rnor_SHR_Utx7134,200,027 - 134,209,527 (+)NCBIUTH_Rnor_SHR_UtxUTH_Rnor_SHR_Utx
UTH_Rnor_SHRSP_BbbUtx_1.07136,429,543 - 136,439,043 (+)NCBIUTH_Rnor_SHRSP_BbbUtx_1.0UTH_Rnor_SHRSP_BbbUtx_1.0
UTH_Rnor_WKY_Bbb_1.07136,339,980 - 136,348,905 (+)NCBIUTH_Rnor_WKY_Bbb_1.0UTH_Rnor_WKY_Bbb_1.0
Rnor_6.07142,929,381 - 142,938,559 (+)NCBIRnor_6.0Rnor_6.0rn6Rnor6.0
Rnor_6.0 Ensembl7142,929,400 - 142,938,554 (+)Ensemblrn6Rnor6.0
Rnor_5.07140,730,251 - 140,739,416 (+)NCBIRnor_5.0Rnor_5.0rn5
RGSC_v3.47140,029,813 - 140,038,925 (+)NCBIRGSC_v3.4RGSC_v3.4rn4
Celera7128,863,838 - 128,872,961 (+)NCBICelera
RGSC_v3.17140,106,249 - 140,115,362 (+)NCBI
Cytogenetic Map7q36NCBI
ATG101
(Homo sapiens - human)
Human AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
GRCh381252,065,300 - 52,077,495 (+)NCBIGRCh38GRCh38hg38GRCh38
GRCh38.p14 Ensembl1252,069,246 - 52,077,495 (+)Ensemblhg38GRCh38
GRCh371252,463,762 - 52,471,279 (+)NCBIGRCh37GRCh37hg19GRCh37
Build 361250,750,025 - 50,757,546 (+)NCBIBuild 36Build 36hg18NCBI36
Celera1251,266,432 - 51,273,953 (+)NCBICelera
Cytogenetic Map12q13.13NCBI
HuRef1249,496,921 - 49,504,446 (+)NCBIHuRef
CHM1_11252,430,002 - 52,437,523 (+)NCBICHM1_1
T2T-CHM13v2.01252,028,879 - 52,041,075 (+)NCBIT2T-CHM13v2.0
Atg101
(Mus musculus - house mouse)
Mouse AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
GRCm3915101,182,182 - 101,188,855 (+)NCBIGRCm39GRCm39mm39
GRCm39 Ensembl15101,182,153 - 101,188,826 (+)EnsemblGRCm39 EnsemblGRCm39
GRCm3815101,284,301 - 101,290,947 (+)NCBIGRCm38GRCm38mm10GRCm38
GRCm38.p6 Ensembl15101,284,272 - 101,290,945 (+)Ensemblmm10GRCm38
MGSCv3715101,114,732 - 101,121,365 (+)NCBIMGSCv37MGSCv37mm9NCBIm37
MGSCv3615101,112,335 - 101,118,968 (+)NCBIMGSCv36mm8
Celera15103,432,360 - 103,438,993 (+)NCBICelera
Cytogenetic Map15F1NCBI
cM Map1556.67NCBI
Atg101
(Chinchilla lanigera - long-tailed chinchilla)
Chinchilla AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
ChiLan1.0 EnsemblNW_0049555472,436,895 - 2,446,849 (+)Ensembl
ChiLan1.0NW_0049555472,436,974 - 2,446,849 (+)NCBIChiLan1.0ChiLan1.0
ATG101
(Pan paniscus - bonobo/pygmy chimpanzee)
Bonobo AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
NHGRI_mPanPan1-v21042,118,333 - 42,126,618 (-)NCBINHGRI_mPanPan1-v2
NHGRI_mPanPan11242,115,096 - 42,123,381 (-)NCBINHGRI_mPanPan1
Mhudiblu_PPA_v01236,685,897 - 36,693,613 (-)NCBIMhudiblu_PPA_v0Mhudiblu_PPA_v0panPan3
PanPan1.11237,462,533 - 37,470,277 (-)NCBIPanPan1.1PanPan1.1panPan2
PanPan1.1 Ensembl1237,462,533 - 37,470,062 (-)EnsemblpanPan2panpan1.1
ATG101
(Canis lupus familiaris - dog)
Dog AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
CanFam3.1272,931,595 - 2,934,914 (-)NCBICanFam3.1CanFam3.1canFam3CanFam3.1
CanFam3.1 Ensembl272,931,893 - 2,935,123 (-)EnsemblcanFam3CanFam3.1
Dog10K_Boxer_Tasha2743,317,316 - 43,324,391 (+)NCBIDog10K_Boxer_TashaDog10K_Boxer_Tasha
ROS_Cfam_1.0272,928,791 - 2,935,881 (-)NCBIROS_Cfam_1.0ROS_Cfam_1.0
ROS_Cfam_1.0 Ensembl272,928,794 - 2,935,861 (-)EnsemblROS_Cfam_1.0 Ensembl
UMICH_Zoey_3.1272,945,425 - 2,950,072 (-)NCBIUMICH_Zoey_3.1
UNSW_CanFamBas_1.0272,931,788 - 2,938,961 (-)NCBIUNSW_CanFamBas_1.0
UU_Cfam_GSD_1.02743,715,530 - 43,722,606 (+)NCBIUU_Cfam_GSD_1.0UU_Cfam_GSD_1.0
Atg101
(Ictidomys tridecemlineatus - thirteen-lined ground squirrel)
Squirrel AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
HiC_Itri_2NW_02440494563,638,844 - 63,646,743 (-)NCBIHiC_Itri_2
SpeTri2.0 EnsemblNW_0049365129,518,074 - 9,526,961 (+)EnsemblSpeTri2.0 Ensembl
SpeTri2.0NW_0049365129,518,128 - 9,525,986 (+)NCBISpeTri2.0SpeTri2.0SpeTri2.0
ATG101
(Sus scrofa - pig)
Pig AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
Sscrofa11.1 Ensembl517,418,121 - 17,425,624 (+)EnsemblsusScr11Sscrofa11.1
Sscrofa11.1517,418,085 - 17,425,770 (+)NCBISscrofa11.1Sscrofa11.1susScr11Sscrofa11.1
ATG101
(Chlorocebus sabaeus - green monkey)
Green Monkey AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
ChlSab1.11148,243,200 - 48,251,129 (+)NCBIChlSab1.1ChlSab1.1chlSab2
ChlSab1.1 Ensembl1148,243,216 - 48,251,371 (+)EnsemblChlSab1.1 EnsemblchlSab2Vervet-AGM
Vero_WHO_p1.0NW_023666037197,841,049 - 197,849,553 (-)NCBIVero_WHO_p1.0Vero_WHO_p1.0
Atg101
(Heterocephalus glaber - naked mole-rat)
Naked Mole-Rat AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
HetGla_female_1.0 EnsemblNW_004624816255,432 - 262,151 (-)EnsemblHetGla_female_1.0 EnsemblhetGla2HetGla_female_1.0 Ensembl
HetGla 1.0NW_004624816255,443 - 261,636 (-)NCBIHetGla 1.0HetGla 1.0hetGla2
Atg101
(Rattus rattus - black rat)
Black Rat AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
Rrattus_CSIRO_v11261,799,183 - 261,805,547 (-)NCBIRrattus_CSIRO_v1

Variants

.
Variants in Atg101
22 total Variants
miRNA Target Status (No longer updated)

Predicted Target Of
Summary Value
Count of predictions:265
Count of miRNA genes:174
Interacting mature miRNAs:204
Transcripts:ENSRNOT00000010194
Prediction methods:Microtar, Miranda, Rnahybrid, Targetscan
Result types:miRGate_prediction

The detailed report is available here: Full Report CSV TAB Printer

miRNA Target Status data imported from miRGate (http://mirgate.bioinfo.cnio.es/).
For more information about miRGate, see PMID:25858286 or access the full paper here.


QTLs in Region (GRCr8)
The following QTLs overlap with this region.    Full Report CSV TAB Printer Gviewer
RGD IDSymbolNameLODP ValueTraitSub TraitChrStartStopSpecies
1331748Bp215Blood pressure QTL 2154.043arterial blood pressure trait (VT:2000000)mean arterial blood pressure (CMO:0000009)7110725024135371468Rat
1354582Stl11Serum triglyceride level QTL 113.42blood triglyceride amount (VT:0002644)serum triglyceride level (CMO:0000360)7121393075137014596Rat
634331Pia17Pristane induced arthritis QTL 174.7joint integrity trait (VT:0010548)arthritic paw count (CMO:0001460)7132099939137014596Rat
731176Glom5Glomerulus QTL 52.50.0035kidney glomerulus morphology trait (VT:0005325)count of superficial glomeruli not directly contacting the kidney surface (CMO:0001002)798549867137014596Rat
1300112Bp183Blood pressure QTL 1833.51arterial blood pressure trait (VT:2000000)blood pressure time series experimental set point of the baroreceptor response (CMO:0002593)7119280025135560836Rat
61428Scl3Serum cholesterol level QTL 33.2blood cholesterol amount (VT:0000180)serum total cholesterol level (CMO:0000363)7100548246137014596Rat
61397Bw17Body weight QTL 176.3body mass (VT:0001259)body weight (CMO:0000012)795485047137014596Rat
1331746Kidm9Kidney mass QTL 93.934kidney mass (VT:0002707)right kidney wet weight (CMO:0000082)782111245134948181Rat
1358914Bp266Blood pressure QTL 266arterial blood pressure trait (VT:2000000)mean arterial blood pressure (CMO:0000009)7115313999136544683Rat
1331731Bp216Blood pressure QTL 2162.851arterial blood pressure trait (VT:2000000)mean arterial blood pressure (CMO:0000009)7104186212135371468Rat
1549899Stresp8Stress response QTL 84.370.0008stress-related behavior trait (VT:0010451)defensive burying duration (CMO:0001961)792362341137014596Rat
1358891Bp265Blood pressure QTL 2652.21arterial blood pressure trait (VT:2000000)mean arterial blood pressure (CMO:0000009)771827901136544683Rat

Markers in Region
RH142906  
Rat AssemblyChrPosition (strand)SourceJBrowse
mRatBN7.27132,407,514 - 132,407,617 (+)MAPPERmRatBN7.2
Rnor_6.07142,938,431 - 142,938,533NCBIRnor6.0
Rnor_5.07140,739,288 - 140,739,390UniSTSRnor5.0
RGSC_v3.47140,038,805 - 140,038,907UniSTSRGSC3.4
Celera7128,872,833 - 128,872,935UniSTS
RH 3.4 Map71075.3UniSTS
Cytogenetic Map7q36UniSTS
RH139413  
Rat AssemblyChrPosition (strand)SourceJBrowse
mRatBN7.27132,407,219 - 132,407,456 (+)MAPPERmRatBN7.2
Rnor_6.07142,938,136 - 142,938,372NCBIRnor6.0
Rnor_5.07140,738,993 - 140,739,229UniSTSRnor5.0
RGSC_v3.47140,038,510 - 140,038,746UniSTSRGSC3.4
Celera7128,872,538 - 128,872,774UniSTS
RH 3.4 Map71063.4UniSTS
Cytogenetic Map7q36UniSTS


Expression

RNA-SEQ Expression

alimentary part of gastrointestinal system
appendage
circulatory system
ectoderm
endocrine system
endoderm
exocrine system
hemolymphoid system
hepatobiliary system
integumental system
mesenchyme
mesoderm
musculoskeletal system
nervous system
renal system
reproductive system
respiratory system
sensory system
visual system
16 12 67 163 91 90 59 92 59 6 356 192 11 142 81 92 31 17 17

Sequence


Ensembl Acc Id: ENSRNOT00000010194   ⟹   ENSRNOP00000010195
Type: CODING
Position:
Rat AssemblyChrPosition (strand)Source
GRCr8 Ensembl7134,277,237 - 134,286,379 (+)Ensembl
mRatBN7.2 Ensembl7132,398,483 - 132,407,633 (+)Ensembl
Rnor_6.0 Ensembl7142,929,439 - 142,938,551 (+)Ensembl
Ensembl Acc Id: ENSRNOT00000087415   ⟹   ENSRNOP00000071030
Type: CODING
Position:
Rat AssemblyChrPosition (strand)Source
GRCr8 Ensembl7134,277,237 - 134,286,113 (+)Ensembl
mRatBN7.2 Ensembl7132,398,493 - 132,407,595 (+)Ensembl
Rnor_6.0 Ensembl7142,929,400 - 142,938,554 (+)Ensembl
Ensembl Acc Id: ENSRNOT00000161382   ⟹   ENSRNOP00000109584
Type: CODING
Position:
Rat AssemblyChrPosition (strand)Source
GRCr8 Ensembl7134,272,847 - 134,286,379 (+)Ensembl
RefSeq Acc Id: NM_001007659   ⟹   NP_001007660
RefSeq Status: VALIDATED
Type: CODING
Position:
Rat AssemblyChrPosition (strand)Source
GRCr87134,277,245 - 134,286,387 (+)NCBI
mRatBN7.27132,398,501 - 132,407,643 (+)NCBI
Rnor_6.07142,929,417 - 142,938,559 (+)NCBI
Rnor_5.07140,730,251 - 140,739,416 (+)NCBI
RGSC_v3.47140,029,813 - 140,038,925 (+)RGD
Celera7128,863,838 - 128,872,961 (+)NCBI
Sequence:
RefSeq Acc Id: NM_001271114   ⟹   NP_001258043
RefSeq Status: VALIDATED
Type: CODING
Position:
Rat AssemblyChrPosition (strand)Source
GRCr87134,277,245 - 134,286,387 (+)NCBI
mRatBN7.27132,398,501 - 132,407,643 (+)NCBI
Rnor_6.07142,929,417 - 142,938,559 (+)NCBI
Rnor_5.07140,730,251 - 140,739,416 (+)NCBI
Celera7128,863,838 - 128,872,961 (+)NCBI
Sequence:
RefSeq Acc Id: XM_006242324   ⟹   XP_006242386
Type: CODING
Position:
Rat AssemblyChrPosition (strand)Source
GRCr87134,277,223 - 134,286,379 (+)NCBI
mRatBN7.27132,398,477 - 132,407,636 (+)NCBI
Rnor_6.07142,929,381 - 142,938,556 (+)NCBI
Rnor_5.07140,730,251 - 140,739,416 (+)NCBI
Sequence:
Protein Sequences
Protein RefSeqs NP_001007660 (Get FASTA)   NCBI Sequence Viewer  
  NP_001258043 (Get FASTA)   NCBI Sequence Viewer  
  XP_006242386 (Get FASTA)   NCBI Sequence Viewer  
GenBank Protein AAH79170 (Get FASTA)   NCBI Sequence Viewer  
  EDL86901 (Get FASTA)   NCBI Sequence Viewer  
  EDL86902 (Get FASTA)   NCBI Sequence Viewer  
Ensembl Protein ENSRNOP00000010195
GenBank Protein Q6AY69 (Get FASTA)   NCBI Sequence Viewer  
RefSeq Acc Id: NP_001007660   ⟸   NM_001007659
- Peptide Label: isoform 1
- UniProtKB: A0A8L2Q495 (UniProtKB/TrEMBL),   A0A8L2R4A7 (UniProtKB/TrEMBL)
- Sequence:
RefSeq Acc Id: NP_001258043   ⟸   NM_001271114
- Peptide Label: isoform 2
- UniProtKB: Q6AY69 (UniProtKB/Swiss-Prot),   A6KCP1 (UniProtKB/TrEMBL),   A0A8L2R4A7 (UniProtKB/TrEMBL)
- Sequence:
RefSeq Acc Id: XP_006242386   ⟸   XM_006242324
- Peptide Label: isoform X1
- UniProtKB: Q6AY69 (UniProtKB/Swiss-Prot),   A6KCP1 (UniProtKB/TrEMBL),   A0A8L2R4A7 (UniProtKB/TrEMBL)
- Sequence:
Ensembl Acc Id: ENSRNOP00000071030   ⟸   ENSRNOT00000087415
Ensembl Acc Id: ENSRNOP00000010195   ⟸   ENSRNOT00000010194
Ensembl Acc Id: ENSRNOP00000109584   ⟸   ENSRNOT00000161382

Protein Structures
Name Modeler Protein Id AA Range Protein Structure
AF-Q6AY69-F1-model_v2 AlphaFold Q6AY69 1-218 view protein structure

Transcriptome

eQTL   View at Phenogen
WGCNA   View at Phenogen
Tissue/Strain Expression   View at Phenogen

Promoters
RGD ID:13695668
Promoter ID:EPDNEW_R6193
Type:multiple initiation site
Name:Atg101_1
Description:autophagy related 101
SO ACC ID:SO:0000170
Source:EPDNEW (Eukaryotic Promoter Database, http://epd.vital-it.ch/)
Experiment Methods:Single-end sequencing.
Position:
Rat AssemblyChrPosition (strand)Source
Rnor_6.07142,929,416 - 142,929,476EPDNEW

Additional Information

Database Acc Id Source(s)
AGR Gene RGD:1359310 AgrOrtholog
BioCyc Gene G2FUF-31992 BioCyc
Ensembl Genes ENSRNOG00000007756 Ensembl, ENTREZGENE
Ensembl Transcript ENSRNOT00000010194 ENTREZGENE
IMAGE_CLONE IMAGE:7129890 IMAGE-MGC_LOAD
InterPro ATG101 UniProtKB/Swiss-Prot
KEGG Report rno:300240 UniProtKB/Swiss-Prot
MGC_CLONE MGC:94201 IMAGE-MGC_LOAD
NCBI Gene 300240 ENTREZGENE
PANTHER AUTOPHAGY-RELATED PROTEIN 101 UniProtKB/Swiss-Prot
  PTHR13292 UniProtKB/Swiss-Prot
Pfam ATG101 UniProtKB/Swiss-Prot
PhenoGen Atg101 PhenoGen
RatGTEx ENSRNOG00000007756 RatGTEx
UniProt A0A8L2Q495 ENTREZGENE, UniProtKB/TrEMBL
  A0A8L2R4A7 ENTREZGENE, UniProtKB/TrEMBL
  A0ABK0LFL9_RAT UniProtKB/TrEMBL
  A6KCP1 ENTREZGENE, UniProtKB/TrEMBL
  ATGA1_RAT UniProtKB/Swiss-Prot, ENTREZGENE


Nomenclature History
Date Current Symbol Current Name Previous Symbol Previous Name Description Reference Status
2014-11-13 Atg101  autophagy related 101  RGD1359310  similar to RIKEN cDNA 9430023L20  Nomenclature updated to reflect human and mouse nomenclature 1299863 APPROVED
2006-03-30 RGD1359310  similar to RIKEN cDNA 9430023L20  MGC94201    Symbol updated 1299863 APPROVED
2005-07-29 MGC94201  similar to RIKEN cDNA 9430023L20      Symbol and Name status set to provisional 70820 PROVISIONAL