DDX39B (DExD-box helicase 39B) - Rat Genome Database

Name: DDX39B
Creator: Rat Genome Database
License: Creative Commons CC BY 4.0
Keywords: Rat Gene RGD Genome

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Gene: DDX39B (DExD-box helicase 39B) Chlorocebus sabaeus

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Symbol:

DDX39B

Name:

DExD-box helicase 39B

RGD ID:

18616637

Description:

ENCODES a protein that exhibits ATP hydrolysis activity (ortholog); protein-containing complex binding (ortholog); INVOLVED IN liver development (ortholog); positive regulation of cell growth involved in cardiac muscle cell development (ortholog); positive regulation of DNA biosynthetic process (ortholog); PARTICIPATES IN NXF1-NXT1 export pathway; spliceosome pathway; ASSOCIATED WITH megacolon (ortholog); Neurodevelopmental Disorders (ortholog); proteasome-associated autoinflammatory syndrome 1 (ortholog); FOUND IN nuclear matrix (ortholog); protein-containing complex (ortholog)

Type:

protein-coding

RefSeq Status:

MODEL

Previously known as:

DEAD-box helicase 39B; spliceosome RNA helicase DDX39B

RGD Orthologs

Alliance Orthologs

More Info

more info ...

More Info

Species	Gene symbol and name	Data Source	Assertion derived from	less info ...
Orthologs ¹				less info ...
Homo sapiens (human):	DDX39B (DExD-box helicase 39B)	NCBI	Ortholog
Mus musculus (house mouse):	Ddx39b (DEAD box helicase 39b)	Transitive Ortholog Pipeline	Transitive Ortholog Pipeline
Rattus norvegicus (Norway rat):	Ddx39b (DExD-box helicase 39B)	Transitive Ortholog Pipeline	Transitive Ortholog Pipeline
Chinchilla lanigera (long-tailed chinchilla):	Ddx39b (DExD-box helicase 39B)	Transitive Ortholog Pipeline	Transitive Ortholog Pipeline
Pan paniscus (bonobo/pygmy chimpanzee):	DDX39B (DExD-box helicase 39B)	Transitive Ortholog Pipeline	Transitive Ortholog Pipeline
Canis lupus familiaris (dog):	DDX39B (DExD-box helicase 39B)	Transitive Ortholog Pipeline	Transitive Ortholog Pipeline
Ictidomys tridecemlineatus (thirteen-lined ground squirrel):	Ddx39b (DExD-box helicase 39B)	Transitive Ortholog Pipeline	Transitive Ortholog Pipeline
Sus scrofa (pig):	DDX39B (DExD-box helicase 39B)	Transitive Ortholog Pipeline	Transitive Ortholog Pipeline
Heterocephalus glaber (naked mole-rat):	Ddx39b (DExD-box helicase 39B)	Transitive Ortholog Pipeline	Transitive Ortholog Pipeline

Latest Assembly:

ChlSab1.1 - Green Monkey 1.1 Assembly

Position:

Green Monkey Assembly	Chr	Position (strand)	Source	Genome Browsers
Green Monkey Assembly	Chr	Position (strand)	Source	JBrowse	NCBI	UCSC	Ensembl
ChlSab1.1	17	40,445,258 - 40,457,379 (+)	NCBI	ChlSab1.1	ChlSab1.1	chlSab2
ChlSab1.1 Ensembl	17	40,447,061 - 40,456,826 (+)	Ensembl	ChlSab1.1	ChlSab1.1 Ensembl	chlSab2
Vero_WHO_p1.0	NW_023666044	31,430,956 - 31,442,672 (-)	NCBI	Vero_WHO_p1.0	Vero_WHO_p1.0

JBrowse:

View Region in Genome Browser (JBrowse)

Model

Annotation Click to see Annotation Detail View

Disease Annotations Click to see Annotation Detail View

megacolon (ISO)

Neurodevelopmental Disorders (ISO)

proteasome-associated autoinflammatory syndrome 1 (ISO)

type 1 diabetes mellitus (ISO)

Gene Ontology Annotations Click to see Annotation Detail View

Biological Process

liver development (ISO)

mRNA export from nucleus (IEA)

mRNA processing (IEA)

mRNA splicing, via spliceosome (IEA)

mRNA transport (IEA)

positive regulation of cell growth involved in cardiac muscle cell development (ISO)

positive regulation of DNA biosynthetic process (ISO)

positive regulation of translation (ISO)

positive regulation of vascular associated smooth muscle cell proliferation (ISO)

RNA secondary structure unwinding (IEA)

RNA splicing (IEA)

spliceosomal complex assembly (IEA)

Cellular Component

cytoplasm (IEA)

nuclear matrix (ISO)

nuclear speck (IEA)

nucleus (IEA)

protein-containing complex (ISO)

spliceosomal complex (IEA)

transcription export complex (IEA)

U4 snRNP (IEA)

U6 snRNP (IEA)

Molecular Function

ATP binding (IEA)

ATP hydrolysis activity (ISO)

ATP-dependent activity, acting on RNA (IEA)

ATP-dependent protein binding (IEA)

helicase activity (IEA)

hydrolase activity (IEA)

identical protein binding (IEA)

nucleic acid binding (IEA)

protein binding (ISO)

protein-containing complex binding (ISO)

RNA helicase activity (IEA)

U4 snRNA binding (IEA)

U6 snRNA binding (IEA)

Molecular Pathway Annotations Click to see Annotation Detail View

NXF1-NXT1 export pathway (ISO)

spliceosome pathway (ISO)

References

Additional References at PubMed

PMID:22301074	PMID:30032202

Genomics

Comparative Map Data

DDX39B
(Chlorocebus sabaeus - green monkey)

Green Monkey Assembly	Chr	Position (strand)	Source	Genome Browsers
Green Monkey Assembly	Chr	Position (strand)	Source	JBrowse	NCBI	UCSC	Ensembl
ChlSab1.1	17	40,445,258 - 40,457,379 (+)	NCBI	ChlSab1.1	ChlSab1.1	chlSab2
ChlSab1.1 Ensembl	17	40,447,061 - 40,456,826 (+)	Ensembl	ChlSab1.1	ChlSab1.1 Ensembl	chlSab2
Vero_WHO_p1.0	NW_023666044	31,430,956 - 31,442,672 (-)	NCBI	Vero_WHO_p1.0	Vero_WHO_p1.0

DDX39B
(Homo sapiens - human)

Human Assembly	Chr	Position (strand)	Source	Genome Browsers
Human Assembly	Chr	Position (strand)	Source	JBrowse	NCBI	UCSC	Ensembl
GRCh38	6	31,530,226 - 31,542,003 (-)	NCBI	GRCh38	GRCh38	hg38	GRCh38
GRCh38.p14 Ensembl	6	31,530,219 - 31,542,448 (-)	Ensembl	GRCh38		hg38	GRCh38
GRCh37	6	31,498,003 - 31,509,780 (-)	NCBI	GRCh37	GRCh37	hg19	GRCh37
Build 36	6	31,605,975 - 31,618,204 (-)	NCBI	NCBI36	Build 36	hg18	NCBI36
Build 34	6	31,605,974 - 31,617,904	NCBI
Celera	6	33,096,234 - 33,108,459 (-)	NCBI		Celera
Cytogenetic Map	6	p21.33	NCBI
HuRef	6	31,284,705 - 31,296,997 (-)	NCBI		HuRef
CHM1_1	6	31,500,137 - 31,512,390 (-)	NCBI		CHM1_1
T2T-CHM13v2.0	6	31,383,322 - 31,395,095 (-)	NCBI		T2T-CHM13v2.0

Ddx39b
(Mus musculus - house mouse)

Mouse Assembly	Chr	Position (strand)	Source	Genome Browsers
Mouse Assembly	Chr	Position (strand)	Source	JBrowse	NCBI	UCSC	Ensembl
GRCm39	17	35,460,722 - 35,472,683 (+)	NCBI	GRCm39	GRCm39	mm39
GRCm39 Ensembl	17	35,460,722 - 35,472,683 (+)	Ensembl		GRCm39 Ensembl
GRCm38	17	35,241,746 - 35,253,707 (+)	NCBI	GRCm38	GRCm38	mm10	GRCm38
GRCm38.p6 Ensembl	17	35,241,746 - 35,253,707 (+)	Ensembl	GRCm38		mm10	GRCm38
MGSCv37	17	35,378,691 - 35,390,652 (+)	NCBI	GRCm37	MGSCv37	mm9	NCBIm37
MGSCv36	17	34,355,767 - 34,367,554 (+)	NCBI		MGSCv36	mm8
MGSCv36	17	34,849,922 - 34,861,736 (+)	NCBI		MGSCv36	mm8
Celera	17	38,343,889 - 38,346,696 (+)	NCBI		Celera
Cytogenetic Map	17	B1	NCBI
cM Map	17	18.6	NCBI

Ddx39b
(Rattus norvegicus - Norway rat)

Rat Assembly	Chr	Position (strand)	Source	Genome Browsers
Rat Assembly	Chr	Position (strand)	Source	JBrowse	NCBI	UCSC	Ensembl
GRCr8	20	3,577,200 - 3,589,651 (-)	NCBI		GRCr8
mRatBN7.2	20	3,547,702 - 3,585,064 (-)	NCBI	mRatBN7.2	mRatBN7.2
mRatBN7.2 Ensembl	20	3,572,056 - 3,584,996 (-)	Ensembl		mRatBN7.2 Ensembl
UTH_Rnor_SHR_Utx	20	4,272,550 - 4,284,961 (-)	NCBI	Rnor_SHR	UTH_Rnor_SHR_Utx
UTH_Rnor_SHRSP_BbbUtx_1.0	20	3,634,613 - 3,647,024 (-)	NCBI	Rnor_SHRSP	UTH_Rnor_SHRSP_BbbUtx_1.0
UTH_Rnor_WKY_Bbb_1.0	20	4,172,085 - 4,184,507 (-)	NCBI	Rnor_WKY	UTH_Rnor_WKY_Bbb_1.0
Rnor_6.0	20	4,806,103 - 4,818,600 (-)	NCBI	Rnor6.0	Rnor_6.0	rn6	Rnor6.0
Rnor_6.0 Ensembl	20	4,806,108 - 4,818,568 (-)	Ensembl	Rnor6.0		rn6	Rnor6.0
Rnor_5.0	20	6,886,503 - 6,898,979 (-)	NCBI	Rnor5.0	Rnor_5.0	rn5	Rnor5.0
RGSC_v3.4	20	3,611,128 - 3,623,578 (-)	NCBI	RGSC3.4	RGSC_v3.4	rn4	RGSC3.4
RGSC_v3.1	20	3,611,355 - 3,623,754 (-)	NCBI
Celera	20	4,440,470 - 4,452,912 (+)	NCBI		Celera
Cytogenetic Map	20	p12	NCBI

Ddx39b
(Chinchilla lanigera - long-tailed chinchilla)

Chinchilla Assembly	Chr	Position (strand)	Source	Genome Browsers
Chinchilla Assembly	Chr	Position (strand)	Source	JBrowse	NCBI	UCSC	Ensembl
ChiLan1.0 Ensembl	NW_004955437	72,835 - 84,384 (-)	Ensembl	ChiLan1.0
ChiLan1.0	NW_004955437	72,835 - 84,045 (-)	NCBI	ChiLan1.0	ChiLan1.0

DDX39B
(Pan paniscus - bonobo/pygmy chimpanzee)

Bonobo Assembly	Chr	Position (strand)	Source	Genome Browsers
Bonobo Assembly	Chr	Position (strand)	Source	JBrowse	NCBI	UCSC	Ensembl
NHGRI_mPanPan1-v2	5	46,007,204 - 46,019,088 (-)	NCBI		NHGRI_mPanPan1-v2
NHGRI_mPanPan1	6	41,968,724 - 41,980,648 (-)	NCBI		NHGRI_mPanPan1
Mhudiblu_PPA_v0	6	31,191,333 - 31,203,246 (-)	NCBI	Mhudiblu_PPA_v0	Mhudiblu_PPA_v0	panPan3
PanPan1.1	6	32,081,210 - 32,093,135 (-)	NCBI	panpan1.1	PanPan1.1	panPan2
PanPan1.1 Ensembl	6	32,081,210 - 32,092,569 (-)	Ensembl	panpan1.1		panPan2

DDX39B
(Canis lupus familiaris - dog)

Dog Assembly	Chr	Position (strand)	Source	Genome Browsers
Dog Assembly	Chr	Position (strand)	Source	JBrowse	NCBI	UCSC	Ensembl
CanFam3.1	12	1,032,377 - 1,044,465 (-)	NCBI	CanFam3.1	CanFam3.1	canFam3	CanFam3.1
CanFam3.1 Ensembl	12	1,032,368 - 1,044,492 (-)	Ensembl	CanFam3.1		canFam3	CanFam3.1
Dog10K_Boxer_Tasha	12	1,169,048 - 1,179,440 (-)	NCBI		Dog10K_Boxer_Tasha
ROS_Cfam_1.0	12	1,177,608 - 1,189,705 (-)	NCBI		ROS_Cfam_1.0
ROS_Cfam_1.0 Ensembl	12	1,176,147 - 1,190,041 (-)	Ensembl		ROS_Cfam_1.0 Ensembl
UMICH_Zoey_3.1	12	1,036,642 - 1,047,031 (-)	NCBI		UMICH_Zoey_3.1
UNSW_CanFamBas_1.0	12	1,104,617 - 1,114,937 (-)	NCBI		UNSW_CanFamBas_1.0
UU_Cfam_GSD_1.0	12	1,171,204 - 1,181,685 (-)	NCBI		UU_Cfam_GSD_1.0

Ddx39b
(Ictidomys tridecemlineatus - thirteen-lined ground squirrel)

Squirrel Assembly	Chr	Position (strand)	Source	Genome Browsers
Squirrel Assembly	Chr	Position (strand)	Source	JBrowse	NCBI	UCSC	Ensembl
HiC_Itri_2	NW_024404946	35,580,562 - 35,597,123 (-)	NCBI		HiC_Itri_2
SpeTri2.0 Ensembl	NW_004936727	1,968,094 - 1,980,785 (+)	Ensembl	SpeTri2.0	SpeTri2.0 Ensembl
SpeTri2.0	NW_004936727	1,968,032 - 1,980,789 (+)	NCBI	SpeTri2.0	SpeTri2.0		SpeTri2.0

DDX39B
(Sus scrofa - pig)

Pig Assembly	Chr	Position (strand)	Source	Genome Browsers
Pig Assembly	Chr	Position (strand)	Source	JBrowse	NCBI	UCSC	Ensembl
Sscrofa11.1 Ensembl	7	23,658,088 - 23,670,031 (-)	Ensembl	Sscrofa11.1		susScr11	Sscrofa11.1
Sscrofa11.1	7	23,658,089 - 23,669,095 (-)	NCBI	Sscrofa11.1	Sscrofa11.1	susScr11	Sscrofa11.1
Sscrofa10.2	7	27,582,945 - 27,593,980 (+)	NCBI	Sscrofa10.2	Sscrofa10.2	susScr3

Ddx39b
(Heterocephalus glaber - naked mole-rat)

Naked Mole-Rat Assembly	Chr	Position (strand)	Source	Genome Browsers
Naked Mole-Rat Assembly	Chr	Position (strand)	Source	JBrowse	NCBI	UCSC	Ensembl
HetGla_female_1.0 Ensembl	NW_004624754	24,657,914 - 24,667,579 (+)	Ensembl	HetGla_female_1.0	HetGla_female_1.0 Ensembl	hetGla2
HetGla 1.0	NW_004624754	24,657,019 - 24,667,225 (+)	NCBI	HetGla_female_1.0	HetGla 1.0	hetGla2

Expression

RNA-SEQ Expression

Click on a value in the shaded box below the category label to view a detailed expression data table for that system.

ectoderm	endocrine system	hemolymphoid system	mesenchyme	mesoderm	nervous system	reproductive system
76	36	20	36	36	60	20

Sequence

Nucleotide Sequences


RefSeq Transcripts	XM_007973138	(Get FASTA)	NCBI Sequence Viewer	Search GEO for Microarray Profiles
	XM_007973139	(Get FASTA)	NCBI Sequence Viewer	Search GEO for Microarray Profiles
GenBank Nucleotide	AC241846	(Get FASTA)	NCBI Sequence Viewer	Search GEO for Microarray Profiles
	JACDXN010000012	(Get FASTA)	NCBI Sequence Viewer	Search GEO for Microarray Profiles

Ensembl Acc Id:

ENSCSAT00000006567 ⟹ ENSCSAP00000004768

Type:

CODING

Position:

Green Monkey Assembly	Chr	Position (strand)	Source
ChlSab1.1 Ensembl	17	40,447,061 - 40,456,826 (+)	Ensembl

RefSeq Acc Id:

XM_007973138 ⟹ XP_007971329

Type:

CODING

Position:

Green Monkey Assembly	Chr	Position (strand)	Source
ChlSab1.1	17	40,445,258 - 40,457,379 (+)	NCBI
Vero_WHO_p1.0	NW_023666044	31,430,956 - 31,442,672 (-)	NCBI

Sequence:

show sequence

CCCTCAGCTAGGGCCTGCCGTTTCCTAGTGCTTGCCCAGCCGCCCTCAGGTCACCTTCACTACC
GGGCCAAGGACCCCGTGGGAACTCGCAGCCCTCGCCACACCGGTTCCTCGCACATCCATGGAGT
GCCTAGAGAGAAGACCTGCATGGCAAAAACCTAAACGAAGAGATGAGAGGCATGGAGAGGAGTA
CGACAAGAGAAGAAAGATAACAGTGGGAGGAGACGTTAGTTTCCTTTATACCTTTTGTTACTGG
CGGTAGCAGGGAAGTTAGAAACGGTTTTAAAACAAATTTCAGACAGGCATTTTCCAAAGGCAAG
CCTGGAGCGCACGGATCTGTACAACCGCGGAAGGCCCCGTTTCCGGTCCCTTGCGCCTGCGCTC
TTGCAGCCAAGAAGGCGGGAGGCTGGAGTAGAGAGAAGCCTGCAACCGGAAGTGAAGGCAGATT
TCCCTCCTTCGTCGCTGTTGCTGCCGCCATACGCGCTCTGCCTGCTTAGCTCTTCTGTTAGAAA
TAGTATCTTTATTTTCCTTTGCTGTTCCTCAATCCCCTACTCTTCACCCCTTGTTTTCACCTGT
TTTGCGAGAACCCATCCAGATCCCCCTTCCCGTCTTCCCCTGCCGGCCCAGTTATGGCAGAGAA
CGATGTGGACAATGAGCTCTTGGACTATGAAGATGATGAGGTGGAGACAGCAGCTGGGGGAGAT
GGGGCTGAGGCCCCTGCCAAGAAGGATGTCAAGGGCTCCTATGTCTCCATCCACAGCTCTGGCT
TTCGTGACTTCCTGCTCAAGCCAGAGTTGCTCCGGGCCATTGTCGACTGTGGCTTTGAGCATCC
GTCAGAAGTCCAGCATGAGTGCATCCCTCAGGCCATTCTGGGAATGGATGTCCTGTGCCAGGCC
AAGTCAGGCATGGGAAAGACAGCAGTGTTTGTGTTGGCCACACTGCAACAGCTGGAGCCAGTTA
CTGGGCAGGTGTCTGTGCTGGTGATGTGTCACACTCGGGAGTTGGCTTTTCAGATCAGCAAGGA
ATATGAGCGCTTCTCTAAATACATGCCCAATGTCAAGGTTGCTGTTTTTTTTGGTGGTCTGTCT
ATCAAGAAGGATGAAGAGGTGCTGAAGAAGAACTGCCCGCATATCGTCGTGGGGACTCCAGGCC
GTATCCTAGCCCTGGCTCGAAATAAGAGCCTCAACCTCAAACACATTAAACACTTTATTTTGGA
TGAATGTGATAAGATGCTTGAACAGCTCGACATGCGTCGGGATGTCCAGGAAATTTTTCGCATG
ACCCCCCACGAGAAGCAGGTCATGATGTTCAGTGCTACCTTGAGCAAAGAGATCCGTCCAGTCT
GCCGCAAGTTCATGCAAGATCCAATGGAGATCTTCGTGGATGATGAGACGAAGTTGACGCTGCA
TGGGTTGCAGCAGTACTACGTGAAACTGAAGGACAACGAGAAGAACCGGAAACTCTTTGACCTT
CTGGATGTCCTTGAGTTCAACCAGGTGGTGATCTTTGTGAAGTCTGTGCAGCGGTGCATTGCCT
TGGCCCAGCTACTGGTGGAGCAGAACTTTCCAGCCATTGCCATCCACCGTGGGATGCCCCAGGA
GGAGAGGCTTTCTCGGTATCAGCAGTTTAAAGATTTTCAACGACGAATTCTTGTGGCTACCAAC
CTATTTGGCCGAGGCATGGACATCGAGCGGGTGAACATTGCTTTTAATTATGACATGCCTGAGG
ATTCTGACACCTACCTACATCGGGTGGCCAGAGCAGGCCGGTTTGGCACCAAGGGCTTGGCTAT
CACATTTGTGTCCGATGAGAATGATGCCAAGATCCTCAATGATGTGCAGGATCGCTTTGAGGTC
AATATTAGTGAGCTGCCTGATGAGATAGACATCTCCTCCTACATTGAACAGACACGGTAGAAGA
CTCGCCCATTCTGGAATGTGACCGTCTGTCCTTCAGGAGAGGACACCAGGGTGGGGGTGAAGGA
GACACTACTGCCCCCACCCCTGACAGCCCCCACCCCATGGCTTCCATCTTTTGCATCACCACCA
CTCCTGAACCCCCATTTCTGATTTGTCAGAATTTTTTTTAACAAAACTAAAAATGAAACACATG
TGTCTGTGGTATCTATAA

hide sequence

Protein Sequences


Protein RefSeqs	XP_007971329	(Get FASTA)	NCBI Sequence Viewer
Ensembl Protein	ENSCSAP00000004768.1

RefSeq Acc Id:	XP_007971329 ⟸ XM_007973138
- UniProtKB:	A0A0D9R829 (UniProtKB/TrEMBL)

Ensembl Acc Id:

ENSCSAP00000004768 ⟸ ENSCSAT00000006567

Additional Information

External Database Links

Database	Acc Id	Source(s)
Ensembl Genes	ENSCSAG00000008517	Ensembl, ENTREZGENE, UniProtKB/TrEMBL
Ensembl Transcript	ENSCSAT00000006567.1	UniProtKB/TrEMBL
Gene3D-CATH	3.40.50.300	UniProtKB/TrEMBL
InterPro	DEAD/DEAH_box_helicase_dom	UniProtKB/TrEMBL
	Helicase_ATP-bd	UniProtKB/TrEMBL
	Helicase_C	UniProtKB/TrEMBL
	P-loop_NTPase	UniProtKB/TrEMBL
	RNA_helicase_DEAD_Q_motif	UniProtKB/TrEMBL
NCBI Gene	DDX39B	ENTREZGENE
PANTHER	ATP-DEPENDENT RNA HELICASE DBP3	UniProtKB/TrEMBL
	SPLICEOSOME RNA HELICASE DDX39B	UniProtKB/TrEMBL
Pfam	DEAD	UniProtKB/TrEMBL
	Helicase_C	UniProtKB/TrEMBL
PROSITE	HELICASE_ATP_BIND_1	UniProtKB/TrEMBL
	HELICASE_CTER	UniProtKB/TrEMBL
	Q_MOTIF	UniProtKB/TrEMBL
SMART	DEXDc	UniProtKB/TrEMBL
	HELICc	UniProtKB/TrEMBL
Superfamily-SCOP	SSF52540	UniProtKB/TrEMBL
UniProt	A0A0D9R829	ENTREZGENE, UniProtKB/TrEMBL

Gene Annotator (Functional Annotation) unavailable	Gene Annotator (Annotation Distribution) unavailable	Variant Visualizer (Genomic Variants) unavailble Variant Visualizer (Genomic Variants) unavailable
Gene Annotator (Functional Annotation)	Gene Annotator (Annotation Distribution)	Variant Visualizer (Genomic Variants)

InterViewer (Protein-Protein Interactions) unavailable	Gviewer (Genome Viewer) unavailable	Variant Visualizer (Damaging Variants) unavailble Variant Visualizer (Damaging Variants) unavailable
InterViewer (Protein-Protein Interactions)	GViewer (Genome Viewer)	Variant Visualizer (Damaging Variants)

Gene Annotator (Annotation Comparison) unavailable	OLGA (Gene List Generator) unavailable
Gene Annotator (Annotation Comparison)	OLGA (Gene List Generator)	Excel (Download)

MOET (Multi-Ontology Enrichement) unavailable	GOLF (Gene-Ortholog Location Finder) unavailable
MOET (Multi-Ontology Enrichement)	GOLF (Gene-Ortholog Location Finder)

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