B3gnt4 (UDP-GlcNAc:betaGal beta-1,3-N-acetylglucosaminyltransferase 4) - Rat Genome Database

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Gene: B3gnt4 (UDP-GlcNAc:betaGal beta-1,3-N-acetylglucosaminyltransferase 4) Mus musculus
Analyze
Symbol: B3gnt4
Name: UDP-GlcNAc:betaGal beta-1,3-N-acetylglucosaminyltransferase 4
RGD ID: 1320044
MGI Page MGI
Description: Predicted to enable N-acetyllactosaminide beta-1,3-N-acetylglucosaminyltransferase activity. Predicted to be involved in poly-N-acetyllactosamine biosynthetic process and protein O-linked glycosylation. Predicted to be located in Golgi apparatus and membrane. Predicted to be active in Golgi membrane. Is expressed in embryo. Orthologous to human B3GNT4 (UDP-GlcNAc:betaGal beta-1,3-N-acetylglucosaminyltransferase 4).
Type: protein-coding
RefSeq Status: VALIDATED
Previously known as: 1010001G17Rik; beta-1,3-Gn-T4; beta-1,3-N-acetylglucosaminyltransferase 4; beta3Gn-T4; BGnT-4; N-acetyllactosaminide beta-1,3-N-acetylglucosaminyltransferase 4
RGD Orthologs
Human
Rat
Chinchilla
Bonobo
Dog
Squirrel
Pig
Green Monkey
Naked Mole-Rat
Alliance Genes
More Info more info ...
Latest Assembly: GRCm39 - Mouse Genome Assembly GRCm39
Position:
Mouse AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
GRCm395123,646,519 - 123,649,945 (+)NCBIGRCm39GRCm39mm39
GRCm39 Ensembl5123,648,523 - 123,649,945 (+)EnsemblGRCm39 Ensembl
GRCm385123,508,461 - 123,511,882 (+)NCBIGRCm38GRCm38mm10GRCm38
GRCm38.p6 Ensembl5123,510,460 - 123,511,882 (+)EnsemblGRCm38mm10GRCm38
MGSCv375123,960,469 - 123,961,891 (+)NCBIGRCm37MGSCv37mm9NCBIm37
MGSCv365123,771,078 - 123,772,500 (+)NCBIMGSCv36mm8
Celera5120,597,308 - 120,598,730 (+)NCBICelera
Cytogenetic Map5FNCBI
cM Map563.02NCBI
JBrowse: View Region in Genome Browser (JBrowse)
Model


Disease Annotations     Click to see Annotation Detail View

Gene-Chemical Interaction Annotations     Click to see Annotation Detail View
Gene Ontology Annotations     Click to see Annotation Detail View

Biological Process

Cellular Component

Molecular Pathway Annotations     Click to see Annotation Detail View
References

References - curated
# Reference Title Reference Citation
1. Functional annotation of a full-length mouse cDNA collection. Kawai J, etal., Nature. 2001 Feb 8;409(6821):685-90.
2. MGDs mouse GO annotations MGD data from the GO Consortium
3. Analysis of the mouse transcriptome based on functional annotation of 60,770 full-length cDNAs. Okazaki Y, etal., Nature. 2002 Dec 5;420(6915):563-73.
4. KEGG Annotation Import Pipeline Pipeline to import KEGG annotations from KEGG into RGD
5. Data Import for Chemical-Gene Interactions RGD automated import pipeline for gene-chemical interactions
Additional References at PubMed
PMID:12477932   PMID:14681479   PMID:20178780   PMID:21267068   PMID:21873635  


Genomics

Comparative Map Data
B3gnt4
(Mus musculus - house mouse)
Mouse AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
GRCm395123,646,519 - 123,649,945 (+)NCBIGRCm39GRCm39mm39
GRCm39 Ensembl5123,648,523 - 123,649,945 (+)EnsemblGRCm39 Ensembl
GRCm385123,508,461 - 123,511,882 (+)NCBIGRCm38GRCm38mm10GRCm38
GRCm38.p6 Ensembl5123,510,460 - 123,511,882 (+)EnsemblGRCm38mm10GRCm38
MGSCv375123,960,469 - 123,961,891 (+)NCBIGRCm37MGSCv37mm9NCBIm37
MGSCv365123,771,078 - 123,772,500 (+)NCBIMGSCv36mm8
Celera5120,597,308 - 120,598,730 (+)NCBICelera
Cytogenetic Map5FNCBI
cM Map563.02NCBI
B3GNT4
(Homo sapiens - human)
Human AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
GRCh3812122,203,709 - 122,208,952 (+)NCBIGRCh38GRCh38hg38GRCh38
GRCh38.p14 Ensembl12122,203,681 - 122,208,952 (+)EnsemblGRCh38hg38GRCh38
GRCh3712122,688,256 - 122,693,499 (+)NCBIGRCh37GRCh37hg19GRCh37
Build 3612121,254,181 - 121,258,037 (+)NCBINCBI36Build 36hg18NCBI36
Build 3412121,213,107 - 121,216,963NCBI
Celera12122,277,574 - 122,281,430 (+)NCBICelera
Cytogenetic Map12q24.31NCBI
HuRef12119,649,164 - 119,653,020 (+)NCBIHuRef
CHM1_112122,656,521 - 122,660,380 (+)NCBICHM1_1
T2T-CHM13v2.012122,199,497 - 122,204,740 (+)NCBIT2T-CHM13v2.0
B3gnt4
(Rattus norvegicus - Norway rat)
Rat AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
GRCr81238,729,653 - 38,734,911 (-)NCBIGRCr8
mRatBN7.21233,070,221 - 33,073,904 (-)NCBImRatBN7.2mRatBN7.2
mRatBN7.2 Ensembl1233,060,416 - 33,073,854 (-)EnsemblmRatBN7.2 Ensembl
UTH_Rnor_SHR_Utx1234,249,189 - 34,250,602 (-)NCBIRnor_SHRUTH_Rnor_SHR_Utx
UTH_Rnor_SHRSP_BbbUtx_1.01234,860,540 - 34,861,953 (-)NCBIRnor_SHRSPUTH_Rnor_SHRSP_BbbUtx_1.0
UTH_Rnor_WKY_Bbb_1.01233,912,429 - 33,913,842 (-)NCBIRnor_WKYUTH_Rnor_WKY_Bbb_1.0
Rnor_6.01238,503,361 - 38,506,793 (-)NCBIRnor6.0Rnor_6.0rn6Rnor6.0
Rnor_6.0 Ensembl1238,503,361 - 38,504,774 (-)EnsemblRnor6.0rn6Rnor6.0
Rnor_5.01240,384,776 - 40,388,648 (-)NCBIRnor5.0Rnor_5.0rn5Rnor5.0
RGSC_v3.41234,209,650 - 34,211,063 (-)NCBIRGSC3.4RGSC_v3.4rn4RGSC3.4
RGSC_v3.11234,073,292 - 34,074,342 (-)NCBI
Celera1234,755,422 - 34,756,835 (-)NCBICelera
Cytogenetic Map12q16NCBI
B3gnt4
(Chinchilla lanigera - long-tailed chinchilla)
Chinchilla AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
ChiLan1.0NW_0049554826,359,967 - 6,365,344 (-)NCBIChiLan1.0ChiLan1.0
B3GNT4
(Pan paniscus - bonobo/pygmy chimpanzee)
Bonobo AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
NHGRI_mPanPan1-v210130,294,008 - 130,297,877 (+)NCBINHGRI_mPanPan1-v2
NHGRI_mPanPan112130,293,033 - 130,294,253 (+)NCBINHGRI_mPanPan1
Mhudiblu_PPA_v012119,804,502 - 119,814,294 (+)NCBIMhudiblu_PPA_v0Mhudiblu_PPA_v0panPan3
PanPan1.112123,172,431 - 123,181,427 (+)NCBIpanpan1.1PanPan1.1panPan2
PanPan1.1 Ensembl12123,172,431 - 123,176,283 (+)Ensemblpanpan1.1panPan2
B3GNT4
(Canis lupus familiaris - dog)
Dog AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
CanFam3.1267,145,120 - 7,147,732 (-)NCBICanFam3.1CanFam3.1canFam3CanFam3.1
CanFam3.1 Ensembl267,145,262 - 7,146,335 (-)EnsemblCanFam3.1canFam3CanFam3.1
Dog10K_Boxer_Tasha267,305,308 - 7,308,871 (-)NCBIDog10K_Boxer_Tasha
ROS_Cfam_1.0267,396,617 - 7,400,186 (-)NCBIROS_Cfam_1.0
ROS_Cfam_1.0 Ensembl267,396,760 - 7,400,411 (-)EnsemblROS_Cfam_1.0 Ensembl
UMICH_Zoey_3.1267,328,767 - 7,332,313 (-)NCBIUMICH_Zoey_3.1
UNSW_CanFamBas_1.0267,419,678 - 7,423,246 (-)NCBIUNSW_CanFamBas_1.0
UU_Cfam_GSD_1.0267,375,419 - 7,378,983 (-)NCBIUU_Cfam_GSD_1.0
B3gnt4
(Ictidomys tridecemlineatus - thirteen-lined ground squirrel)
Squirrel AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
HiC_Itri_2NW_024405118157,206,764 - 157,211,525 (+)NCBIHiC_Itri_2
SpeTri2.0 EnsemblNW_0049365582,269,367 - 2,270,413 (-)EnsemblSpeTri2.0SpeTri2.0 Ensembl
SpeTri2.0NW_0049365582,269,182 - 2,272,664 (-)NCBISpeTri2.0SpeTri2.0SpeTri2.0
B3GNT4
(Sus scrofa - pig)
Pig AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
Sscrofa11.1 Ensembl1430,520,077 - 30,523,060 (-)EnsemblSscrofa11.1susScr11Sscrofa11.1
Sscrofa11.11430,520,075 - 30,524,275 (-)NCBISscrofa11.1Sscrofa11.1susScr11Sscrofa11.1
Sscrofa10.21432,327,298 - 32,337,894 (+)NCBISscrofa10.2Sscrofa10.2susScr3
B3GNT4
(Chlorocebus sabaeus - green monkey)
Green Monkey AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
ChlSab1.111117,552,133 - 117,555,112 (+)NCBIChlSab1.1ChlSab1.1chlSab2
Vero_WHO_p1.0NW_023666037127,771,241 - 127,774,996 (-)NCBIVero_WHO_p1.0Vero_WHO_p1.0
B3gnt4
(Heterocephalus glaber - naked mole-rat)
Naked Mole-Rat AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
HetGla 1.0NW_00462474722,440,284 - 22,441,539 (+)NCBIHetGla_female_1.0HetGla 1.0hetGla2

Variants

.
Variants in B3gnt4
132 total Variants
miRNA Target Status

Predicted Target Of
Summary Value
Count of predictions:170
Count of miRNA genes:159
Interacting mature miRNAs:168
Transcripts:ENSMUST00000031384
Prediction methods:Miranda, Rnahybrid, Targetscan
Result types:miRGate_prediction

The detailed report is available here: Full Report CSV TAB Printer

miRNA Target Status data imported from miRGate (http://mirgate.bioinfo.cnio.es/).
For more information about miRGate, see PMID:25858286 or access the full paper here.


QTLs in Region (GRCm39)
The following QTLs overlap with this region.    Full Report CSV TAB Printer Gviewer
RGD IDSymbolNameLODP ValueTraitSub TraitChrStartStopSpecies
11341716Rvfs3_mRift Valley fever susceptibility 3 (mouse)526441234138455402Mouse
4142064Tmc1m4_mTmc1 modifier 4 (mouse)Not determined554500177127001072Mouse
26884395Humsd2_mhumerus midshaft diameter 2, 10 week (mouse)570057343132528839Mouse
27226719Tibw6_mtibia width 6, proximal, 16 week (mouse)572857343139985755Mouse
25314315Mlh1fc4_mMLH1 foci count 4 (mouse)574460661132528839Mouse
13506928Recrq8_mrecombination rate in male meiosis QTL 8 (mouse)574460661132528839Mouse
1300968Skts4_mskin tumor susceptibility 4 (mouse)Not determined589418876124238239Mouse
1301362Prnr1_mprion resistance 1 (mouse)Not determined589418876146787935Mouse
11049562Lmr24f_mleishmaniasis resistance 24f (mouse)590320962124321041Mouse
1301457Cfsw2_mcystic fibrosis survival to weaning 2 (mouse)Not determined590561006124561151Mouse
1300993Hycdc_mhypercapnic duty cycle (mouse)Not determined590561006124561151Mouse
9587780Afw16_mabdominal fat weight QTL 16 (mouse)Not determined592830166126830285Mouse
1301974Chab7_mcholesterol absorption 7 (mouse)Not determined595233214129233330Mouse
1357644Egrd1_mearly growth rate, direct effect 1 (mouse)Not determined595434580129434786Mouse
10402486Dipa1_mdrug induced psychomotor activation 1 (mouse)Not determined595465549129465722Mouse
13464244Ahl19_mage related hearing loss, early onset 19 (mouse)595465549129465722Mouse
1558774Lith17_mlithogenic gene 17 (mouse)Not determined595465549129465722Mouse
1300940Actd3_mactivity-distance traveled 3 (mouse)Not determined596320385130320533Mouse
1300659Pcd8ts1_mp-glycoprotein positive CD8 T cell subset 1 (mouse)Not determined597021753131021986Mouse
1301858Smdq1_msegregation of mitochondrial DNA QTL 1 (mouse)Not determined597021753131021986Mouse
4141519Hmtb5_mhemostasis and thrombosis bleeding time 5 (mouse)Not determined99949422126882944Mouse
12904945Tammq2_mtibialis anterior muscle mass QTL 2 (mouse)5100665965134665965Mouse
12904959Smmq1_msoleus muscle mass QTL 1 (mouse)5100665965134665965Mouse
12859290Criq2_mCitrobacter rodentium infection QTL 2 (mouse)5101332579135332579Mouse
1301624Aevm2_mautoimmune extremity vasculitis in MRL mice 2 (mouse)Not determined5101863832135864028Mouse
14746986Manh58_mmandible shape 58 (mouse)5102446283136446283Mouse
11532736Sluc33b_msusceptibility to lung cancer 33b (mouse)5102945037136945196Mouse
10412202Bbaa26_mB.burgdorferi-associated arthritis 26 (mouse)Not determined5104583688138555455Mouse
10412239Alpq7_malcohol preference QTL 7 (mouse)Not determined5104705939138705939Mouse
10043890Trigq4_mtriglyceride QTL 4 (mouse)Not determined5107906299141906412Mouse
4142473Chlq11_mcirculating hormone level QTL 11 (mouse)Not determined5107906299141906412Mouse
13464243Ahl20_mage related hearing loss, early onset 20 (mouse)5107906299141906412Mouse
4141139Hmtb4_mhemostasis and thrombosis rebleeding time 4 (mouse)Not determined108762509126882944Mouse
10412197Bbaa25_mB.burgdorferi-associated arthritis 25 (mouse)Not determined5109167240143167381Mouse
13464251Ahl21_mage related hearing loss, early onset 21 (mouse)5109167240143167381Mouse
1301543Hypch_mhypercholesterolemia (mouse)Not determined5109238624139118905Mouse
1300913Bwefm_mbody weight females and males day 10 (mouse)Not determined5109439348143439459Mouse
1301102Cia14_mcollagen induced arthritis 14 (mouse)Not determined5113320385138555455Mouse
1357662Lprq4_mlipoprotein QTL 4 (mouse)Not determined5114021753129632606Mouse
1301226Bbaa2_mB.burgdorferi-associated arthritis 2 (mouse)Not determined5115156883149157015Mouse
1300827Cora1_mcorrelation in cytokine production 1 (mouse)Not determined5115156883149157015Mouse
1301857Bglq14_mbody growth late QTL 14 (mouse)Not determined5118948964151758149Mouse
10412191Bbaa24_mB.burgdorferi-associated arthritis 24 (mouse)Not determined5119945037141700466Mouse
4141081Nidd7k_mNidd7 on KK-A (mouse)Not determined119945037149347326Mouse
11528552Scram1_mspinal cord resistance to astrocytoma modifier 1 (mouse)5121455402151758149Mouse
1301986Bpq4_mblood pressure QTL 4 (mouse)Not determined5121555236151758149Mouse

Markers in Region
AU040403  
Mouse AssemblyChrPosition (strand)SourceJBrowse
GRCm385123,509,863 - 123,510,133UniSTSGRCm38
MGSCv375123,959,872 - 123,960,142UniSTSGRCm37
Celera5120,596,701 - 120,596,981UniSTS
Cytogenetic Map5FUniSTS
Whitehead/MRC_RH51501.06UniSTS
UniSTS:235312  
Mouse AssemblyChrPosition (strand)SourceJBrowse
GRCm385123,512,155 - 123,512,262UniSTSGRCm38
MGSCv375123,962,164 - 123,962,271UniSTSGRCm37
Celera5120,599,003 - 120,599,110UniSTS
Cytogenetic Map5FUniSTS
UniSTS:487717  
Mouse AssemblyChrPosition (strand)SourceJBrowse
GRCm385123,510,505 - 123,511,879UniSTSGRCm38
MGSCv375123,960,514 - 123,961,888UniSTSGRCm37
Celera5120,597,353 - 120,598,727UniSTS
Cytogenetic Map5FUniSTS
UniSTS:493301  
Mouse AssemblyChrPosition (strand)SourceJBrowse
GRCm385123,510,500 - 123,511,879UniSTSGRCm38
MGSCv375123,960,509 - 123,961,888UniSTSGRCm37
Celera5120,597,348 - 120,598,727UniSTS
Cytogenetic Map5FUniSTS


Expression


Sequence


Reference Sequences
RefSeq Acc Id: ENSMUST00000031384   ⟹   ENSMUSP00000031384
Type: CODING
Position:
Mouse AssemblyChrPosition (strand)Source
GRCm39 Ensembl5123,648,523 - 123,649,945 (+)Ensembl
GRCm38.p6 Ensembl5123,510,460 - 123,511,882 (+)Ensembl
RefSeq Acc Id: NM_001382854   ⟹   NP_001369783
RefSeq Status: VALIDATED
Type: CODING
Position:
Mouse AssemblyChrPosition (strand)Source
GRCm395123,646,545 - 123,649,844 (+)NCBI
GRCm385123,508,482 - 123,511,781 (+)NCBI
Sequence:
RefSeq Acc Id: NM_198611   ⟹   NP_941013
RefSeq Status: VALIDATED
Type: CODING
Position:
Mouse AssemblyChrPosition (strand)Source
GRCm395123,648,523 - 123,649,945 (+)NCBI
GRCm385123,510,460 - 123,511,882 (+)NCBI
MGSCv375123,960,469 - 123,961,891 (+)RGD
Celera5120,597,308 - 120,598,730 (+)RGD
cM Map5 ENTREZGENE
Sequence:
RefSeq Acc Id: XM_006530313   ⟹   XP_006530376
Type: CODING
Position:
Mouse AssemblyChrPosition (strand)Source
GRCm395123,646,519 - 123,649,839 (+)NCBI
GRCm385123,508,461 - 123,511,776 (+)NCBI
Sequence:
RefSeq Acc Id: XM_011248199   ⟹   XP_011246501
Type: CODING
Position:
Mouse AssemblyChrPosition (strand)Source
GRCm395123,646,903 - 123,649,839 (+)NCBI
GRCm385123,508,840 - 123,511,776 (+)NCBI
Sequence:
RefSeq Acc Id: XM_011248200   ⟹   XP_011246502
Type: CODING
Position:
Mouse AssemblyChrPosition (strand)Source
GRCm395123,646,591 - 123,649,839 (+)NCBI
GRCm385123,508,528 - 123,511,776 (+)NCBI
Sequence:
RefSeq Acc Id: XM_036165093   ⟹   XP_036020986
Type: CODING
Position:
Mouse AssemblyChrPosition (strand)Source
GRCm395123,646,591 - 123,649,839 (+)NCBI
Sequence:
Reference Protein Sequences
RefSeq Acc Id: NP_941013   ⟸   NM_198611
- UniProtKB: E9QPF4 (UniProtKB/Swiss-Prot),   Q923H4 (UniProtKB/Swiss-Prot),   Q1RLK6 (UniProtKB/Swiss-Prot)
- Sequence:
RefSeq Acc Id: XP_006530376   ⟸   XM_006530313
- Peptide Label: isoform X2
- UniProtKB: E9QPF4 (UniProtKB/Swiss-Prot),   Q923H4 (UniProtKB/Swiss-Prot),   Q1RLK6 (UniProtKB/Swiss-Prot)
- Sequence:
RefSeq Acc Id: XP_011246502   ⟸   XM_011248200
- Peptide Label: isoform X2
- UniProtKB: E9QPF4 (UniProtKB/Swiss-Prot),   Q923H4 (UniProtKB/Swiss-Prot),   Q1RLK6 (UniProtKB/Swiss-Prot)
- Sequence:
RefSeq Acc Id: XP_011246501   ⟸   XM_011248199
- Peptide Label: isoform X1
- Sequence:
RefSeq Acc Id: NP_001369783   ⟸   NM_001382854
- Peptide Label: precursor
- UniProtKB: Q1RLK6 (UniProtKB/Swiss-Prot),   E9QPF4 (UniProtKB/Swiss-Prot),   Q923H4 (UniProtKB/Swiss-Prot)
RefSeq Acc Id: ENSMUSP00000031384   ⟸   ENSMUST00000031384
RefSeq Acc Id: XP_036020986   ⟸   XM_036165093
- Peptide Label: isoform X2
- UniProtKB: Q1RLK6 (UniProtKB/Swiss-Prot),   E9QPF4 (UniProtKB/Swiss-Prot),   Q923H4 (UniProtKB/Swiss-Prot)

Protein Structures
Name Modeler Protein Id AA Range Protein Structure
AF-Q1RLK6-F1-model_v2 AlphaFold Q1RLK6 1-350 view protein structure

Promoters
RGD ID:6837149
Promoter ID:MM_KWN:44037
Type:CpG-Island
SO ACC ID:SO:0000170
Source:MPROMDB
Tissues & Cell Lines:BoneMarrow_0Hour,   BoneMarrow_2Hour,   BoneMarrow_4Hour,   Brain,   ES_Cell,   Kidney,   Liver,   Lung,   MEF_B4,   MEF_B6
Transcripts:NM_198611
Position:
Mouse AssemblyChrPosition (strand)Source
MGSCv365123,959,471 - 123,960,817 (+)MPROMDB

Additional Information

Database Acc Id Source(s)
AGR Gene MGI:2680208 AgrOrtholog
Ensembl Genes ENSMUSG00000029431 Ensembl, ENTREZGENE, UniProtKB/Swiss-Prot
Ensembl Transcript ENSMUST00000031384 ENTREZGENE
  ENSMUST00000031384.6 UniProtKB/Swiss-Prot
Gene3D-CATH 3.90.550.50 UniProtKB/Swiss-Prot
InterPro Glyco_trans_31 UniProtKB/Swiss-Prot
KEGG Report mmu:231727 UniProtKB/Swiss-Prot
MGD MGI:2680208 ENTREZGENE
NCBI Gene 231727 ENTREZGENE
PANTHER N-ACETYLLACTOSAMINIDE BETA-1,3-N-ACETYLGLUCOSAMINYLTRANSFERASE 4 UniProtKB/Swiss-Prot
  PTHR11214 UniProtKB/Swiss-Prot
Pfam Galactosyl_T UniProtKB/Swiss-Prot
PhenoGen B3gnt4 PhenoGen
UniProt B3GN4_MOUSE UniProtKB/Swiss-Prot
  E9QPF4 ENTREZGENE
  Q1RLK6 ENTREZGENE
  Q923H4 ENTREZGENE
UniProt Secondary E9QPF4 UniProtKB/Swiss-Prot
  Q923H4 UniProtKB/Swiss-Prot