Tmem126a (transmembrane protein 126A) - Rat Genome Database

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Gene: Tmem126a (transmembrane protein 126A) Mus musculus
Analyze
Symbol: Tmem126a
Name: transmembrane protein 126A
RGD ID: 1313846
MGI Page MGI
Description: Acts upstream of or within toll-like receptor 4 signaling pathway. Located in plasma membrane. Human ortholog(s) of this gene implicated in optic atrophy 7. Orthologous to human TMEM126A (transmembrane protein 126A).
Type: protein-coding
RefSeq Status: VALIDATED
Previously known as: 1810020E01Rik
RGD Orthologs
Human
Rat
Chinchilla
Bonobo
Dog
Squirrel
Pig
Green Monkey
Naked Mole-Rat
Alliance Genes
More Info more info ...
Related Pseudogenes: Gm15186   Gm18330   Gm18388   Gm2477  
Latest Assembly: GRCm39 - Mouse Genome Assembly GRCm39
Position:
Mouse AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
GRCm39790,099,908 - 90,106,411 (-)NCBIGRCm39GRCm39mm39
GRCm39 Ensembl790,099,908 - 90,106,437 (-)EnsemblGRCm39 Ensembl
GRCm38790,450,700 - 90,457,203 (-)NCBIGRCm38GRCm38mm10GRCm38
GRCm38.p6 Ensembl790,450,700 - 90,457,229 (-)EnsemblGRCm38mm10GRCm38
MGSCv37797,599,222 - 97,605,718 (-)NCBIGRCm37MGSCv37mm9NCBIm37
MGSCv36790,325,904 - 90,332,400 (-)NCBIMGSCv36mm8
Celera787,778,196 - 87,784,676 (-)NCBICelera
Cytogenetic Map7E1NCBI
cM Map750.59NCBI
JBrowse: View Region in Genome Browser (JBrowse)
Model


Disease Annotations     Click to see Annotation Detail View

Gene Ontology Annotations     Click to see Annotation Detail View

Cellular Component

Molecular Function

Phenotype Annotations     Click to see Annotation Detail View

Mammalian Phenotype
References

References - curated
# Reference Title Reference Citation
1. Functional annotation of a full-length mouse cDNA collection. Kawai J, etal., Nature. 2001 Feb 8;409(6821):685-90.
2. MGDs mouse GO annotations MGD data from the GO Consortium
3. Analysis of the mouse transcriptome based on functional annotation of 60,770 full-length cDNAs. Okazaki Y, etal., Nature. 2002 Dec 5;420(6915):563-73.
4. OMIM Disease Annotation Pipeline OMIM Disease Annotation Pipeline
5. Mouse MP Annotation Import Pipeline RGD automated import pipeline
6. ClinVar Automated Import and Annotation Pipeline RGD automated import pipeline for ClinVar variants, variant-to-disease annotations and gene-to-disease annotations
7. Data Import for Chemical-Gene Interactions RGD automated import pipeline for gene-chemical interactions
Additional References at PubMed
PMID:8889548   PMID:10349636   PMID:11042159   PMID:11076861   PMID:12477932   PMID:14610273   PMID:15489334   PMID:15782199   PMID:16141072   PMID:16141073   PMID:17967808   PMID:18614015  
PMID:18799693   PMID:19723774   PMID:21267068   PMID:21677750   PMID:21873635   PMID:22885069   PMID:23912999   PMID:25549946   PMID:28378742   PMID:30550785   PMID:30973865  


Genomics

Comparative Map Data
Tmem126a
(Mus musculus - house mouse)
Mouse AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
GRCm39790,099,908 - 90,106,411 (-)NCBIGRCm39GRCm39mm39
GRCm39 Ensembl790,099,908 - 90,106,437 (-)EnsemblGRCm39 Ensembl
GRCm38790,450,700 - 90,457,203 (-)NCBIGRCm38GRCm38mm10GRCm38
GRCm38.p6 Ensembl790,450,700 - 90,457,229 (-)EnsemblGRCm38mm10GRCm38
MGSCv37797,599,222 - 97,605,718 (-)NCBIGRCm37MGSCv37mm9NCBIm37
MGSCv36790,325,904 - 90,332,400 (-)NCBIMGSCv36mm8
Celera787,778,196 - 87,784,676 (-)NCBICelera
Cytogenetic Map7E1NCBI
cM Map750.59NCBI
TMEM126A
(Homo sapiens - human)
Human AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
GRCh381185,647,967 - 85,656,542 (+)NCBIGRCh38GRCh38hg38GRCh38
GRCh38.p14 Ensembl1185,647,967 - 85,656,547 (+)EnsemblGRCh38hg38GRCh38
GRCh371185,359,011 - 85,367,586 (+)NCBIGRCh37GRCh37hg19GRCh37
Build 361185,036,693 - 85,045,235 (+)NCBINCBI36Build 36hg18NCBI36
Celera1182,668,277 - 82,676,912 (+)NCBICelera
Cytogenetic Map11q14.1NCBI
HuRef1181,656,256 - 81,664,891 (+)NCBIHuRef
CHM1_11185,243,754 - 85,252,388 (+)NCBICHM1_1
T2T-CHM13v2.01185,584,644 - 85,593,220 (+)NCBIT2T-CHM13v2.0
Tmem126a
(Rattus norvegicus - Norway rat)
Rat AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
GRCr81153,837,015 - 153,843,192 (-)NCBIGRCr8
mRatBN7.21144,424,481 - 144,430,730 (-)NCBImRatBN7.2mRatBN7.2
mRatBN7.2 Ensembl1144,422,703 - 144,430,628 (-)EnsemblmRatBN7.2 Ensembl
UTH_Rnor_SHR_Utx1152,347,274 - 152,353,446 (-)NCBIRnor_SHRUTH_Rnor_SHR_Utx
UTH_Rnor_SHRSP_BbbUtx_1.01159,528,451 - 159,534,610 (-)NCBIRnor_SHRSPUTH_Rnor_SHRSP_BbbUtx_1.0
UTH_Rnor_WKY_Bbb_1.01152,402,370 - 152,408,529 (-)NCBIRnor_WKYUTH_Rnor_WKY_Bbb_1.0
Rnor_6.01156,283,105 - 156,291,057 (+)NCBIRnor6.0Rnor_6.0rn6Rnor6.0
Rnor_6.0 Ensembl1156,283,128 - 156,291,052 (+)EnsemblRnor6.0rn6Rnor6.0
Rnor_5.01162,529,113 - 162,537,179 (+)NCBIRnor5.0Rnor_5.0rn5Rnor5.0
RGSC_v3.41147,129,330 - 147,137,259 (-)NCBIRGSC3.4RGSC_v3.4rn4RGSC3.4
RGSC_v3.11147,207,735 - 147,215,665 (-)NCBI
Celera1142,643,098 - 142,651,027 (-)NCBICelera
Cytogenetic Map1q32NCBI
Tmem126a
(Chinchilla lanigera - long-tailed chinchilla)
Chinchilla AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
ChiLan1.0 EnsemblNW_0049554147,216,286 - 7,224,475 (-)EnsemblChiLan1.0
ChiLan1.0NW_0049554147,214,408 - 7,224,499 (-)NCBIChiLan1.0ChiLan1.0
TMEM126A
(Pan paniscus - bonobo/pygmy chimpanzee)
Bonobo AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
NHGRI_mPanPan1-v2986,567,317 - 86,579,574 (+)NCBINHGRI_mPanPan1-v2
NHGRI_mPanPan11187,620,767 - 87,630,548 (+)NCBINHGRI_mPanPan1
Mhudiblu_PPA_v01180,703,855 - 80,712,459 (+)NCBIMhudiblu_PPA_v0Mhudiblu_PPA_v0panPan3
PanPan1.11184,312,434 - 84,321,037 (+)NCBIpanpan1.1PanPan1.1panPan2
PanPan1.1 Ensembl1184,312,434 - 84,321,037 (+)Ensemblpanpan1.1panPan2
TMEM126A
(Canis lupus familiaris - dog)
Dog AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
CanFam3.12113,787,074 - 13,795,779 (-)NCBICanFam3.1CanFam3.1canFam3CanFam3.1
CanFam3.1 Ensembl2113,787,123 - 13,795,739 (-)EnsemblCanFam3.1canFam3CanFam3.1
Dog10K_Boxer_Tasha2113,653,725 - 13,662,471 (-)NCBIDog10K_Boxer_Tasha
ROS_Cfam_1.02113,967,872 - 13,976,597 (-)NCBIROS_Cfam_1.0
ROS_Cfam_1.0 Ensembl2113,967,464 - 13,976,562 (-)EnsemblROS_Cfam_1.0 Ensembl
UMICH_Zoey_3.12113,756,370 - 13,765,102 (-)NCBIUMICH_Zoey_3.1
UNSW_CanFamBas_1.02113,828,697 - 13,837,423 (-)NCBIUNSW_CanFamBas_1.0
UU_Cfam_GSD_1.02113,881,469 - 13,890,217 (-)NCBIUU_Cfam_GSD_1.0
Tmem126a
(Ictidomys tridecemlineatus - thirteen-lined ground squirrel)
Squirrel AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
HiC_Itri_2NW_02440494771,938,905 - 71,945,739 (+)NCBIHiC_Itri_2
SpeTri2.0NW_00493649813,538,565 - 13,545,338 (+)NCBISpeTri2.0SpeTri2.0SpeTri2.0
TMEM126A
(Sus scrofa - pig)
Pig AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
Sscrofa11.1 Ensembl919,576,295 - 19,585,277 (+)EnsemblSscrofa11.1susScr11Sscrofa11.1
Sscrofa11.1919,576,241 - 19,585,869 (+)NCBISscrofa11.1Sscrofa11.1susScr11Sscrofa11.1
Sscrofa10.2921,846,389 - 21,855,417 (+)NCBISscrofa10.2Sscrofa10.2susScr3
TMEM126A
(Chlorocebus sabaeus - green monkey)
Green Monkey AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
ChlSab1.1176,888,688 - 76,897,883 (+)NCBIChlSab1.1ChlSab1.1chlSab2
ChlSab1.1 Ensembl176,888,734 - 76,897,841 (+)EnsemblChlSab1.1ChlSab1.1 EnsemblchlSab2
Vero_WHO_p1.0NW_02366604348,824,623 - 48,833,726 (-)NCBIVero_WHO_p1.0Vero_WHO_p1.0
Tmem126a
(Heterocephalus glaber - naked mole-rat)
Naked Mole-Rat AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
HetGla 1.0NW_0046248454,264,497 - 4,272,973 (-)NCBIHetGla_female_1.0HetGla 1.0hetGla2

Variants

.
Variants in Tmem126a
264 total Variants
miRNA Target Status

Predicted Target Of
Summary Value
Count of predictions:241
Count of miRNA genes:206
Interacting mature miRNAs:223
Transcripts:ENSMUST00000032844, ENSMUST00000136652, ENSMUST00000153393
Prediction methods:Miranda, Rnahybrid, Targetscan
Result types:miRGate_prediction

The detailed report is available here: Full Report CSV TAB Printer

miRNA Target Status data imported from miRGate (http://mirgate.bioinfo.cnio.es/).
For more information about miRGate, see PMID:25858286 or access the full paper here.


QTLs in Region (GRCm39)
The following QTLs overlap with this region.    Full Report CSV TAB Printer Gviewer
RGD IDSymbolNameLODP ValueTraitSub TraitChrStartStopSpecies
25314307Mlh1fc2_mMLH1 foci count 2 (mouse)76502999133501729Mouse
12792978Fbmd3_mfemoral bone mineral density 3, females only (mouse)77050288142367832Mouse
10412199Sst2_msusceptibility to tuberculosis 2 (mouse)Not determined718728794119485380Mouse
1301158Eae4_msusceptibility to experimental allergic encephalomyelitis 4 (mouse)Not determined719147398141919804Mouse
26884404Huml1_mhumerus length 1, 5 week (mouse)730199425108999207Mouse
1301514Rigs1_mradiation induced gastroschisis 1 (mouse)Not determined73628001592394346Mouse
1301082Bbaa16_mB.burgdorferi-associated arthritis 16 (mouse)Not determined736280015116416877Mouse
1300791Abbp3_mA/J and C57BL/6 blood pressure 3 (mouse)Not determined739673887103510010Mouse
10043926Bw1n_mbody weight 1 in NSY (mouse)Not determined745161966118693530Mouse
13208555Lgth10_mbody length 10 (mouse)751649748125599172Mouse
27226751Femd5_mfemur midshaft diameter 5, 10 week (mouse)753249748143153737Mouse
25823170Hrsq7_mhost response to SARS QTL 7, log titer (mouse)754819589116822815Mouse
27226778Femd11_mfemur midshaft diameter 11, 16 week (mouse)756949748134001729Mouse
1300952Hcs1_mhepatocarcinogenesis susceptibility 1 (mouse)Not determined75732612191326352Mouse
1301465Egrm3_mearly growth rate (mouse)Not determined75732612191326352Mouse
1301663Skull9_mskull morphology 9 (mouse)Not determined75732612191326352Mouse
26884408Bzwq13_mbi-zygomatic width QTL 13, 16 week (mouse)761749748121399223Mouse
27095916Scvln15_msacral vertebrae length 2, 16 week (mouse)761749748123699223Mouse
25314305Vmm6_mvariable multisystem mineralization 6, eye (mouse)761849748102749207Mouse
1301383Aem2_manti-erythrocyte autoantibody modifier 2 (mouse)Not determined76413542498135653Mouse
26884381Bzwq2_mbi-zygomatic width QTL 2, 5 week (mouse)76524974896949207Mouse
26884440Sklq4_mskull length QTL 4, 5 week (mouse)765249748126499172Mouse
11553865Stmm1c_mskin tumor modifier of MSM 1c (mouse)76578898699788986Mouse
11553866Stmm1d_mskin tumor modifier of MSM 1d (mouse)76578898699788986Mouse
27095937Ulnl7_mulna length 7, 10 week (mouse)76644974898349207Mouse
1301826Pgia3_mproteoglycan induced arthritis 3 (mouse)Not determined767290094101290240Mouse
1301784Ses2_msalmonella enteritidis susceptibility 2 (mouse)Not determined767290094101290240Mouse
1300620Pgia21_mproteoglycan induced arthritis 21 (mouse)Not determined767290094101290240Mouse
26884418Bzwq7_mbi-zygomatic width QTL 7, 10 week (mouse)767849748105449207Mouse
27226715Metcl3_mmetatarsal-calcaneal length 3, 5 week (mouse)767849748119499223Mouse
26884444Sklq8_mskull length QTL 8, 10 week (mouse)767849748124199223Mouse
26884451Sklq14_mskull length QTL 14, 16 week (mouse)767849748124999172Mouse
1300654Bomd1_mbone mineral density 1 (mouse)Not determined768365595102365702Mouse
27095907Scvln4_msacral vertebrae length 2, 5 week (mouse)769949748120499223Mouse
1301155Bpq7_mblood pressure QTL 7 (mouse)Not determined770143137104143379Mouse
10043973Obq29_mobesity QTL 29 (mouse)Not determined772263372106263372Mouse
4142319Powg_mpost-ovarectomy weight gain (mouse)Not determined7285710592582163Mouse
27095910Pglq13_mpelvic girdle length QTL 13, 16 week (mouse)773049748125699172Mouse
27095933Ulnl3_mulna length 3, 5 week (mouse)773149748143153737Mouse
11553869Stmm2b_mskin tumor modifier of MSM 2a (mouse)773725965107725965Mouse
4142212Png1_mPostnatal growth 1 (mouse)Not determined7432612192394346Mouse
27226745Metcl9_mmetatarsal-calcaneal length 9, 10 week (mouse)774849748122199223Mouse
27226781Tibl3_mtibia length 3, 5 week (mouse)774849748122199223Mouse
1301566Adip3_madiposity 3 (mouse)Not determined775394215109394346Mouse
14746989Manh62_mmandible shape 62 (mouse)775742528109742528Mouse
26884398Humsd6_mhumerus midshaft diameter 6, 16 week (mouse)778649748136901729Mouse
10412110Cctq1_mcentral corneal thickness QTL 1 (mouse)Not determined780551681114551791Mouse
4142500Morq1_mmodifier of Rs1 QTL 1 (mouse)Not determined80551681114551791Mouse
27095930Ulnl10_mulna length 10, 16 week (mouse)780849748126699172Mouse
27095923Pglq3_mpelvic girdle length QTL 3, 5 week (mouse)781249748124199223Mouse
27095903Scvln10_msacral vertebrae length 2, 10 week (mouse)781449748126499172Mouse
27226765Tibl18_mtibia length 18, 16 week (mouse)781849748126499172Mouse
27226763Feml20_mfemur length 20, 16 week (mouse)785049208124199223Mouse
1301997Cia7_mcollagen induced arthritis QTL 7 (mouse)Not determined785365595112533819Mouse
27226792Feml7_mfemur length 7, 5 week (mouse)78614920898949207Mouse
1301647Elpt_melevated leptin (mouse)Not determined786500186120500329Mouse
1301188Heal6_mwound healing/regeneration 6 (mouse)Not determined787043932121044066Mouse
1302062Stheal6_msoft tissue heal 6 (mouse)Not determined787043932121044066Mouse
13463473Hbbcq4_mhemoglobin concentration QTL 4 (mouse)787043932121044066Mouse
1301781Capsq3_mcapsaicin sensitivity related QTL 3 (mouse)Not determined787143137119485380Mouse
27226769Tibl12_mtibia length 12, 10 week (mouse)787249208126499172Mouse
27226790Feml15_mfemur length 15, 10 week (mouse)787249208126499172Mouse
13825246Albc2_malbino coat color QTL 2 (mouse)78764920895649207Mouse
27095929Pglq8_mpelvic girdle length QTL 8, 10 week (mouse)787849208108099207Mouse

Markers in Region
RH128303  
Mouse AssemblyChrPosition (strand)SourceJBrowse
GRCm38X159,588,668 - 159,588,847UniSTSGRCm38
MGSCv37X156,026,600 - 156,026,779UniSTSGRCm37
CeleraX142,833,980 - 142,834,159UniSTS
Cytogenetic MapXF4UniSTS
Cytogenetic Map7E1UniSTS


Expression


Sequence


Reference Sequences
RefSeq Acc Id: ENSMUST00000032844   ⟹   ENSMUSP00000032844
Type: CODING
Position:
Mouse AssemblyChrPosition (strand)Source
GRCm39 Ensembl790,099,908 - 90,106,437 (-)Ensembl
GRCm38.p6 Ensembl790,450,700 - 90,457,229 (-)Ensembl
RefSeq Acc Id: ENSMUST00000136652   ⟹   ENSMUSP00000115803
Type: CODING
Position:
Mouse AssemblyChrPosition (strand)Source
GRCm39 Ensembl790,099,908 - 90,106,416 (-)Ensembl
GRCm38.p6 Ensembl790,450,700 - 90,457,208 (-)Ensembl
RefSeq Acc Id: ENSMUST00000153393
Type: CODING
Position:
Mouse AssemblyChrPosition (strand)Source
GRCm39 Ensembl790,104,413 - 90,106,361 (-)Ensembl
GRCm38.p6 Ensembl790,455,205 - 90,457,153 (-)Ensembl
RefSeq Acc Id: ENSMUST00000207159
Type: CODING
Position:
Mouse AssemblyChrPosition (strand)Source
GRCm39 Ensembl790,099,920 - 90,106,310 (-)Ensembl
GRCm38.p6 Ensembl790,450,712 - 90,457,102 (-)Ensembl
RefSeq Acc Id: ENSMUST00000208639
Type: CODING
Position:
Mouse AssemblyChrPosition (strand)Source
GRCm39 Ensembl790,104,664 - 90,106,393 (-)Ensembl
GRCm38.p6 Ensembl790,455,456 - 90,457,185 (-)Ensembl
RefSeq Acc Id: NM_025460   ⟹   NP_079736
RefSeq Status: VALIDATED
Type: CODING
Position:
Mouse AssemblyChrPosition (strand)Source
GRCm39790,099,908 - 90,106,411 (-)NCBI
GRCm38790,450,700 - 90,457,203 (-)NCBI
MGSCv37797,599,222 - 97,605,718 (-)RGD
Celera787,778,196 - 87,784,676 (-)RGD
cM Map7 ENTREZGENE
Sequence:
RefSeq Acc Id: NR_157050
RefSeq Status: VALIDATED
Type: NON-CODING
Position:
Mouse AssemblyChrPosition (strand)Source
GRCm39790,099,908 - 90,106,411 (-)NCBI
GRCm38790,450,700 - 90,457,203 (-)NCBI
Sequence:
Reference Protein Sequences
RefSeq Acc Id: NP_079736   ⟸   NM_025460
- UniProtKB: Q9D8Y1 (UniProtKB/Swiss-Prot)
- Sequence:
RefSeq Acc Id: ENSMUSP00000032844   ⟸   ENSMUST00000032844
RefSeq Acc Id: ENSMUSP00000115803   ⟸   ENSMUST00000136652

Protein Structures
Name Modeler Protein Id AA Range Protein Structure
AF-Q9D8Y1-F1-model_v2 AlphaFold Q9D8Y1 1-196 view protein structure

Promoters
RGD ID:8664751
Promoter ID:EPDNEW_M10406
Type:initiation region
Name:Tmem126a_1
Description:Mus musculus transmembrane protein 126A , mRNA.
SO ACC ID:SO:0000170
Source:EPDNEW (Eukaryotic Promoter Database, http://epd.vital-it.ch/)
Experiment Methods:Single-end sequencing.
Position:
Mouse AssemblyChrPosition (strand)Source
GRCm38790,457,186 - 90,457,246EPDNEW
RGD ID:6842202
Promoter ID:MM_KWN:51518
Type:CpG-Island
SO ACC ID:SO:0000170
Source:MPROMDB
Tissues & Cell Lines:3T3L1_Day0,   3T3L1_Day1,   3T3L1_Day2,   3T3L1_Day3,   3T3L1_Day4,   3T3L1_Day6,   BoneMarrow_0Hour,   BoneMarrow_2Hour,   BoneMarrow_4Hour,   Brain,   ES_Cell,   Kidney,   Liver,   Lung,   MEF_B4,   MEF_B6,   Spleen
Transcripts:OTTMUST00000077392,   OTTMUST00000077393,   OTTMUST00000077394
Position:
Mouse AssemblyChrPosition (strand)Source
MGSCv36797,605,386 - 97,605,952 (-)MPROMDB

Additional Information

Database Acc Id Source(s)
AGR Gene MGI:1913521 AgrOrtholog
Ensembl Genes ENSMUSG00000030615 Ensembl, ENTREZGENE, UniProtKB/Swiss-Prot, UniProtKB/TrEMBL
Ensembl Transcript ENSMUST00000032844 ENTREZGENE
  ENSMUST00000032844.7 UniProtKB/Swiss-Prot
  ENSMUST00000136652 ENTREZGENE
  ENSMUST00000136652.8 UniProtKB/TrEMBL
InterPro TMEM126 UniProtKB/Swiss-Prot, UniProtKB/TrEMBL
KEGG Report mmu:66271 UniProtKB/Swiss-Prot
MGD MGI:1913521 ENTREZGENE
NCBI Gene 66271 ENTREZGENE
PANTHER PTHR16296 UniProtKB/Swiss-Prot, UniProtKB/TrEMBL
  TRANSMEMBRANE PROTEIN 126A UniProtKB/Swiss-Prot, UniProtKB/TrEMBL
Pfam TMEM126 UniProtKB/Swiss-Prot, UniProtKB/TrEMBL
PhenoGen Tmem126a PhenoGen
UniProt B0LAC9_MOUSE UniProtKB/TrEMBL
  D6RG44_MOUSE UniProtKB/TrEMBL
  Q9D8Y1 ENTREZGENE, UniProtKB/Swiss-Prot