Xylb (xylulokinase) - Rat Genome Database

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Pathways
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Gene: Xylb (xylulokinase) Rattus norvegicus
Analyze
Symbol: Xylb
Name: xylulokinase
RGD ID: 1307372
Description: Predicted to enable xylulokinase activity. Predicted to be involved in phosphorylation and xylulose metabolic process. Predicted to be active in cytosol. Orthologous to human XYLB (xylulokinase); PARTICIPATES IN pentose and glucuronate interconversion pathway; INTERACTS WITH (+)-schisandrin B; 2,3,4,7,8-Pentachlorodibenzofuran; 2,3,7,8-tetrachlorodibenzodioxine.
Type: protein-coding
RefSeq Status: PROVISIONAL
Previously known as: LOC316067; MGC124790; xylulokinase homolog (H. influenzae); xylulose kinase
RGD Orthologs
Human
Mouse
Chinchilla
Bonobo
Dog
Squirrel
Pig
Green Monkey
Naked Mole-Rat
Alliance Genes
More Info more info ...
Latest Assembly: mRatBN7.2 - mRatBN7.2 Assembly
Position:
Rat AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
mRatBN7.28119,093,466 - 119,128,858 (+)NCBImRatBN7.2mRatBN7.2
mRatBN7.2 Ensembl8119,096,029 - 119,128,848 (+)EnsemblmRatBN7.2 Ensembl
UTH_Rnor_SHR_Utx8124,678,128 - 124,710,751 (+)NCBIRnor_SHR
UTH_Rnor_SHRSP_BbbUtx_1.08122,877,145 - 122,909,770 (+)NCBIRnor_SHRSP
UTH_Rnor_WKY_Bbb_1.08120,710,647 - 120,743,316 (+)NCBIRnor_WKY
Rnor_6.08128,041,875 - 128,076,951 (+)NCBIRnor6.0Rnor_6.0rn6Rnor6.0
Rnor_6.0 Ensembl8128,044,084 - 128,076,946 (+)EnsemblRnor6.0rn6Rnor6.0
Rnor_5.08127,248,438 - 127,283,712 (+)NCBIRnor5.0Rnor_5.0rn5Rnor5.0
RGSC_v3.48124,321,366 - 124,354,135 (+)NCBIRGSC3.4RGSC_v3.4rn4RGSC3.4
RGSC_v3.18124,340,874 - 124,373,231 (+)NCBI
Celera8118,247,268 - 118,280,101 (+)NCBICelera
Cytogenetic Map8q32NCBI
JBrowse: View Region in Genome Browser (JBrowse)
Model


Disease Annotations     Click to see Annotation Detail View

Gene Ontology Annotations     Click to see Annotation Detail View

Biological Process

Cellular Component
cytosol  (IBA)

Molecular Function
ATP binding  (IEA)
xylulokinase activity  (IBA,IEA,ISO,ISS)

Molecular Pathway Annotations     Click to see Annotation Detail View
References

References - curated
# Reference Title Reference Citation
1. Phylogenetic-based propagation of functional annotations within the Gene Ontology consortium. Gaudet P, etal., Brief Bioinform. 2011 Sep;12(5):449-62. doi: 10.1093/bib/bbr042. Epub 2011 Aug 27.
2. Rat ISS GO annotations from GOA human gene data--August 2006 GOA data from the GO Consortium
3. KEGG Annotation Import Pipeline Pipeline to import KEGG annotations from KEGG into RGD
4. GOA pipeline RGD automated data pipeline
5. Data Import for Chemical-Gene Interactions RGD automated import pipeline for gene-chemical interactions
Additional References at PubMed
PMID:12477932   PMID:15489334   PMID:23179721   PMID:23376485  


Genomics

Comparative Map Data
Xylb
(Rattus norvegicus - Norway rat)
Rat AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
mRatBN7.28119,093,466 - 119,128,858 (+)NCBImRatBN7.2mRatBN7.2
mRatBN7.2 Ensembl8119,096,029 - 119,128,848 (+)EnsemblmRatBN7.2 Ensembl
UTH_Rnor_SHR_Utx8124,678,128 - 124,710,751 (+)NCBIRnor_SHR
UTH_Rnor_SHRSP_BbbUtx_1.08122,877,145 - 122,909,770 (+)NCBIRnor_SHRSP
UTH_Rnor_WKY_Bbb_1.08120,710,647 - 120,743,316 (+)NCBIRnor_WKY
Rnor_6.08128,041,875 - 128,076,951 (+)NCBIRnor6.0Rnor_6.0rn6Rnor6.0
Rnor_6.0 Ensembl8128,044,084 - 128,076,946 (+)EnsemblRnor6.0rn6Rnor6.0
Rnor_5.08127,248,438 - 127,283,712 (+)NCBIRnor5.0Rnor_5.0rn5Rnor5.0
RGSC_v3.48124,321,366 - 124,354,135 (+)NCBIRGSC3.4RGSC_v3.4rn4RGSC3.4
RGSC_v3.18124,340,874 - 124,373,231 (+)NCBI
Celera8118,247,268 - 118,280,101 (+)NCBICelera
Cytogenetic Map8q32NCBI
XYLB
(Homo sapiens - human)
Human AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
GRCh38338,346,785 - 38,453,041 (+)NCBIGRCh38GRCh38hg38GRCh38
GRCh38.p13 Ensembl338,346,760 - 38,421,348 (+)EnsemblGRCh38hg38GRCh38
GRCh37338,388,276 - 38,468,677 (+)NCBIGRCh37GRCh37hg19GRCh37
Build 36338,363,244 - 38,431,471 (+)NCBINCBI36Build 36hg18NCBI36
Build 34338,363,243 - 38,431,471NCBI
Celera338,323,884 - 38,392,347 (+)NCBICelera
Cytogenetic Map3p22.2NCBI
HuRef338,430,197 - 38,498,798 (+)NCBIHuRef
CHM1_1338,339,982 - 38,408,211 (+)NCBICHM1_1
T2T-CHM13v2.0338,352,536 - 38,459,118 (+)NCBIT2T-CHM13v2.0
Xylb
(Mus musculus - house mouse)
Mouse AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
GRCm399119,186,447 - 119,222,863 (+)NCBIGRCm39GRCm39mm39
GRCm39 Ensembl9119,186,447 - 119,222,863 (+)EnsemblGRCm39 Ensembl
GRCm389119,357,381 - 119,393,797 (+)NCBIGRCm38GRCm38mm10GRCm38
GRCm38.p6 Ensembl9119,357,381 - 119,393,797 (+)EnsemblGRCm38mm10GRCm38
MGSCv379119,266,499 - 119,302,438 (+)NCBIGRCm37MGSCv37mm9NCBIm37
MGSCv369119,206,079 - 119,242,018 (+)NCBIMGSCv36mm8
Celera9119,826,564 - 119,863,083 (+)NCBICelera
Cytogenetic Map9F3NCBI
Xylb
(Chinchilla lanigera - long-tailed chinchilla)
Chinchilla AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
ChiLan1.0 EnsemblNW_00495542725,492,505 - 25,528,740 (-)EnsemblChiLan1.0
ChiLan1.0NW_00495542725,492,505 - 25,528,674 (-)NCBIChiLan1.0ChiLan1.0
XYLB
(Pan paniscus - bonobo/pygmy chimpanzee)
Bonobo AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
NHGRI_mPanPan1338,305,249 - 38,386,454 (+)NCBINHGRI_mPanPan1
Mhudiblu_PPA_v0338,244,960 - 38,349,616 (+)NCBIMhudiblu_PPA_v0Mhudiblu_PPA_v0panPan3
PanPan1.1338,526,968 - 38,631,714 (+)NCBIpanpan1.1PanPan1.1panPan2
PanPan1.1 Ensembl338,526,988 - 38,592,086 (+)Ensemblpanpan1.1panPan2
XYLB
(Canis lupus familiaris - dog)
Dog AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
CanFam3.1238,123,041 - 8,171,803 (+)NCBICanFam3.1CanFam3.1canFam3CanFam3.1
CanFam3.1 Ensembl238,122,197 - 8,170,447 (+)EnsemblCanFam3.1canFam3CanFam3.1
Dog10K_Boxer_Tasha238,163,344 - 8,213,617 (+)NCBIDog10K_Boxer_Tasha
ROS_Cfam_1.0238,411,983 - 8,462,247 (+)NCBIROS_Cfam_1.0
ROS_Cfam_1.0 Ensembl238,412,008 - 8,464,550 (+)EnsemblROS_Cfam_1.0 Ensembl
UMICH_Zoey_3.1238,226,162 - 8,276,351 (+)NCBIUMICH_Zoey_3.1
UNSW_CanFamBas_1.0238,366,621 - 8,416,906 (+)NCBIUNSW_CanFamBas_1.0
UU_Cfam_GSD_1.0238,360,020 - 8,410,270 (+)NCBIUU_Cfam_GSD_1.0
Xylb
(Ictidomys tridecemlineatus - thirteen-lined ground squirrel)
Squirrel AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
HiC_Itri_2NW_024405118194,586,571 - 194,632,938 (+)NCBIHiC_Itri_2
SpeTri2.0 EnsemblNW_00493647327,725,937 - 27,767,102 (+)EnsemblSpeTri2.0
SpeTri2.0NW_00493647327,725,955 - 27,767,319 (+)NCBISpeTri2.0SpeTri2.0SpeTri2.0
XYLB
(Sus scrofa - pig)
Pig AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
Sscrofa11.1 Ensembl1323,173,939 - 23,225,951 (+)EnsemblSscrofa11.1susScr11Sscrofa11.1
Sscrofa11.11323,173,855 - 23,225,583 (+)NCBISscrofa11.1Sscrofa11.1susScr11Sscrofa11.1
Sscrofa10.21325,302,487 - 25,350,851 (-)NCBISscrofa10.2Sscrofa10.2susScr3
XYLB
(Chlorocebus sabaeus - green monkey)
Green Monkey AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
ChlSab1.1151,003,809 - 1,070,385 (-)NCBIChlSab1.1ChlSab1.1chlSab2
ChlSab1.1 Ensembl151,005,340 - 1,069,275 (-)EnsemblChlSab1.1ChlSab1.1 EnsemblchlSab2
Vero_WHO_p1.0NW_0236660639,722,166 - 9,803,123 (-)NCBIVero_WHO_p1.0Vero_WHO_p1.0
Xylb
(Heterocephalus glaber - naked mole-rat)
Naked Mole-Rat AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
HetGla_female_1.0 EnsemblNW_0046247312,619,334 - 2,658,967 (-)EnsemblHetGla_female_1.0HetGla_female_1.0 EnsemblhetGla2
HetGla 1.0NW_0046247312,616,924 - 2,658,675 (-)NCBIHetGla_female_1.0HetGla 1.0hetGla2

Variants

.
Variants in Xylb
248 total Variants
miRNA Target Status

Predicted Target Of
Summary Value
Count of predictions:194
Count of miRNA genes:133
Interacting mature miRNAs:156
Transcripts:ENSRNOT00000019106
Prediction methods:Miranda, Rnahybrid, Targetscan
Result types:miRGate_prediction

The detailed report is available here: Full Report CSV TAB Printer

miRNA Target Status data imported from miRGate (http://mirgate.bioinfo.cnio.es/).
For more information about miRGate, see PMID:25858286 or access the full paper here.


QTLs in Region (mRatBN7.2)
The following QTLs overlap with this region.    Full Report CSV TAB Printer Gviewer
RGD IDSymbolNameLODP ValueTraitSub TraitChrStartStopSpecies
1554321Bmd3Bone mineral density QTL 37.90.0001femur mineral mass (VT:0010011)volumetric bone mineral density (CMO:0001553)840952565123900184Rat
2300181Bmd55Bone mineral density QTL 555.70.0001femur mineral mass (VT:0010011)volumetric bone mineral density (CMO:0001553)876468691121468691Rat
61437Cia6Collagen induced arthritis QTL 6joint integrity trait (VT:0010548)joint inflammation composite score (CMO:0000919)882460758122812818Rat
738011Anxrr9Anxiety related response QTL 96.1exploratory behavior trait (VT:0010471)number of entries into a discrete space in an experimental apparatus (CMO:0000960)893535351123900184Rat
1358893Bp263Blood pressure QTL 2635.01arterial blood pressure trait (VT:2000000)mean arterial blood pressure (CMO:0000009)893965141123900184Rat
1358903Bp252Blood pressure QTL 25270.0001arterial blood pressure trait (VT:2000000)mean arterial blood pressure (CMO:0000009)893965141123900184Rat
738014Anxrr15Anxiety related response QTL 153.60.005locomotor behavior trait (VT:0001392)amount of experiment time spent in a discrete space in an experimental apparatus (CMO:0000958)895718998123900184Rat
2300182Bmd56Bone mineral density QTL 565.4femur mineral mass (VT:0010011)volumetric bone mineral density (CMO:0001553)895718998123900184Rat
724539Cm19Cardiac mass QTL 192.6heart mass (VT:0007028)calculated heart weight (CMO:0000073)8100149864120994388Rat

Markers in Region
RH129038  
Rat AssemblyChrPosition (strand)SourceJBrowse
mRatBN7.28119,128,673 - 119,128,828 (+)MAPPERmRatBN7.2
Rnor_6.08128,076,767 - 128,076,921NCBIRnor6.0
Rnor_5.08127,283,528 - 127,283,682UniSTSRnor5.0
RGSC_v3.48124,353,958 - 124,354,112UniSTSRGSC3.4
Celera8118,279,924 - 118,280,078UniSTS
RH 3.4 Map81286.3UniSTS
Cytogenetic Map8q32UniSTS


Expression


RNA-SEQ Expression
High: > 1000 TPM value   Medium: Between 11 and 1000 TPM
Low: Between 0.5 and 10 TPM   Below Cutoff: < 0.5 TPM

alimentary part of gastrointestinal system circulatory system endocrine system exocrine system hemolymphoid system hepatobiliary system integumental system musculoskeletal system nervous system renal system reproductive system respiratory system appendage
High
Medium 3 12 48 41 41 21 35 28 3
Low 31 9 19 8 11 53 13 8 8
Below cutoff

Sequence


Reference Sequences
RefSeq Acc Id: ENSRNOT00000019106   ⟹   ENSRNOP00000019106
RefSeq Status:
Type: CODING
Position:
Rat AssemblyChrPosition (strand)Source
mRatBN7.2 Ensembl8119,096,037 - 119,128,848 (+)Ensembl
Rnor_6.0 Ensembl8128,044,084 - 128,076,946 (+)Ensembl
RefSeq Acc Id: ENSRNOT00000104648   ⟹   ENSRNOP00000077183
RefSeq Status:
Type: CODING
Position:
Rat AssemblyChrPosition (strand)Source
mRatBN7.2 Ensembl8119,096,077 - 119,128,845 (+)Ensembl
RefSeq Acc Id: ENSRNOT00000120248   ⟹   ENSRNOP00000092265
RefSeq Status:
Type: CODING
Position:
Rat AssemblyChrPosition (strand)Source
mRatBN7.2 Ensembl8119,096,029 - 119,128,765 (+)Ensembl
RefSeq Acc Id: NM_001033704   ⟹   NP_001028876
RefSeq Status: PROVISIONAL
Type: CODING
Position:
Rat AssemblyChrPosition (strand)Source
mRatBN7.28119,096,077 - 119,128,851 (+)NCBI
Rnor_6.08128,044,175 - 128,076,944 (+)NCBI
Rnor_5.08127,248,438 - 127,283,712 (+)NCBI
RGSC_v3.48124,321,366 - 124,354,135 (+)RGD
Celera8118,247,268 - 118,280,101 (+)RGD
Sequence:
RefSeq Acc Id: XM_006244096   ⟹   XP_006244158
RefSeq Status:
Type: CODING
Position:
Rat AssemblyChrPosition (strand)Source
mRatBN7.28119,095,995 - 119,128,858 (+)NCBI
Rnor_6.08128,044,093 - 128,075,357 (+)NCBI
Rnor_5.08127,248,438 - 127,283,712 (+)NCBI
Sequence:
RefSeq Acc Id: XM_017595698   ⟹   XP_017451187
RefSeq Status:
Type: CODING
Position:
Rat AssemblyChrPosition (strand)Source
mRatBN7.28119,093,489 - 119,127,465 (+)NCBI
Rnor_6.08128,044,018 - 128,075,558 (+)NCBI
Sequence:
RefSeq Acc Id: XM_017595699   ⟹   XP_017451188
RefSeq Status:
Type: CODING
Position:
Rat AssemblyChrPosition (strand)Source
mRatBN7.28119,093,486 - 119,128,858 (+)NCBI
Rnor_6.08128,044,018 - 128,076,951 (+)NCBI
Sequence:
RefSeq Acc Id: XM_039081640   ⟹   XP_038937568
RefSeq Status:
Type: CODING
Position:
Rat AssemblyChrPosition (strand)Source
mRatBN7.28119,093,499 - 119,128,858 (+)NCBI
RefSeq Acc Id: XM_039081641   ⟹   XP_038937569
RefSeq Status:
Type: CODING
Position:
Rat AssemblyChrPosition (strand)Source
mRatBN7.28119,093,489 - 119,128,858 (+)NCBI
RefSeq Acc Id: XM_039081642   ⟹   XP_038937570
RefSeq Status:
Type: CODING
Position:
Rat AssemblyChrPosition (strand)Source
mRatBN7.28119,093,466 - 119,113,373 (+)NCBI
RefSeq Acc Id: XM_039081643   ⟹   XP_038937571
RefSeq Status:
Type: CODING
Position:
Rat AssemblyChrPosition (strand)Source
mRatBN7.28119,096,088 - 119,128,858 (+)NCBI
RefSeq Acc Id: XM_039081644   ⟹   XP_038937572
RefSeq Status:
Type: CODING
Position:
Rat AssemblyChrPosition (strand)Source
mRatBN7.28119,093,470 - 119,112,697 (+)NCBI
RefSeq Acc Id: XR_356707
RefSeq Status:
Type: NON-CODING
Position:
Rat AssemblyChrPosition (strand)Source
mRatBN7.28119,093,476 - 119,122,731 (+)NCBI
Rnor_6.08128,041,875 - 128,069,759 (+)NCBI
Rnor_5.08127,248,438 - 127,283,712 (+)NCBI
Sequence:
Reference Sequences
RefSeq Acc Id: NP_001028876   ⟸   NM_001033704
- UniProtKB: A0A8I5Y795 (UniProtKB/TrEMBL)
- Sequence:
RefSeq Acc Id: XP_006244158   ⟸   XM_006244096
- Peptide Label: isoform X5
- Sequence:
RefSeq Acc Id: XP_017451188   ⟸   XM_017595699
- Peptide Label: isoform X3
- Sequence:
RefSeq Acc Id: XP_017451187   ⟸   XM_017595698
- Peptide Label: isoform X2
- Sequence:
RefSeq Acc Id: ENSRNOP00000019106   ⟸   ENSRNOT00000019106
RefSeq Acc Id: XP_038937570   ⟸   XM_039081642
- Peptide Label: isoform X6
RefSeq Acc Id: XP_038937572   ⟸   XM_039081644
- Peptide Label: isoform X8
RefSeq Acc Id: XP_038937569   ⟸   XM_039081641
- Peptide Label: isoform X4
RefSeq Acc Id: XP_038937568   ⟸   XM_039081640
- Peptide Label: isoform X1
RefSeq Acc Id: XP_038937571   ⟸   XM_039081643
- Peptide Label: isoform X7
RefSeq Acc Id: ENSRNOP00000092265   ⟸   ENSRNOT00000120248
RefSeq Acc Id: ENSRNOP00000077183   ⟸   ENSRNOT00000104648
Protein Domains
Carbohydrate kinase FGGY C-terminal   Carbohydrate kinase FGGY N-terminal

Protein Structures
Name Modeler Protein Id AA Range Protein Structure
AF-Q3MIF4-F1-model_v2 AlphaFold Q3MIF4 1-536 view protein structure

Transcriptome

eQTL   View at Phenogen
WGCNA   View at Phenogen
Tissue/Strain Expression   View at Phenogen

Promoters
RGD ID:13696371
Promoter ID:EPDNEW_R6896
Type:initiation region
Name:Xylb_1
Description:xylulokinase
SO ACC ID:SO:0000170
Source:EPDNEW (Eukaryotic Promoter Database, http://epd.vital-it.ch/)
Experiment Methods:Single-end sequencing.
Position:
Rat AssemblyChrPosition (strand)Source
Rnor_6.08128,044,118 - 128,044,178EPDNEW

Additional Information

Database Acc Id Source(s)
AGR Gene RGD:1307372 AgrOrtholog
BioCyc Gene G2FUF-28789 BioCyc
Ensembl Genes ENSRNOG00000014168 Ensembl, ENTREZGENE, UniProtKB/TrEMBL
Ensembl Protein ENSRNOP00000019106.8 UniProtKB/TrEMBL
  ENSRNOP00000077183 ENTREZGENE
  ENSRNOP00000077183.1 UniProtKB/TrEMBL
  ENSRNOP00000092265.1 UniProtKB/TrEMBL
Ensembl Transcript ENSRNOT00000019106.8 UniProtKB/TrEMBL
  ENSRNOT00000104648 ENTREZGENE
  ENSRNOT00000104648.1 UniProtKB/TrEMBL
  ENSRNOT00000120248.1 UniProtKB/TrEMBL
Gene3D-CATH 3.30.420.40 UniProtKB/Swiss-Prot, UniProtKB/TrEMBL
IMAGE_CLONE IMAGE:7930657 IMAGE-MGC_LOAD
InterPro ATPase_NBD UniProtKB/Swiss-Prot, UniProtKB/TrEMBL
  Carb_kinase_FGGY UniProtKB/Swiss-Prot, UniProtKB/TrEMBL
  Carb_kinase_FGGY_C UniProtKB/Swiss-Prot, UniProtKB/TrEMBL
  Carb_kinase_FGGY_N UniProtKB/Swiss-Prot, UniProtKB/TrEMBL
  D-XK_euk UniProtKB/Swiss-Prot, UniProtKB/TrEMBL
KEGG Report rno:316067 UniProtKB/Swiss-Prot
MGC_CLONE MGC:124790 IMAGE-MGC_LOAD
NCBI Gene 316067 ENTREZGENE
PANTHER SUGAR KINASE UniProtKB/Swiss-Prot, UniProtKB/TrEMBL
  XYLULOSE KINASE UniProtKB/Swiss-Prot, UniProtKB/TrEMBL
Pfam FGGY_C UniProtKB/Swiss-Prot, UniProtKB/TrEMBL
  FGGY_N UniProtKB/Swiss-Prot, UniProtKB/TrEMBL
PhenoGen Xylb PhenoGen
PIRSF GlpK UniProtKB/Swiss-Prot, UniProtKB/TrEMBL
Superfamily-SCOP SSF53067 UniProtKB/Swiss-Prot, UniProtKB/TrEMBL
UniProt A0A8I5Y795 ENTREZGENE, UniProtKB/TrEMBL
  A0A8I6AH82_RAT UniProtKB/TrEMBL
  G3V7T5_RAT UniProtKB/TrEMBL
  Q3MIF4 ENTREZGENE, UniProtKB/Swiss-Prot


Nomenclature History
Date Current Symbol Current Name Previous Symbol Previous Name Description Reference Status
2016-07-07 Xylb  xylulokinase  Xylb  xylulokinase homolog (H. influenzae)  Nomenclature updated to reflect human and mouse nomenclature 1299863 APPROVED
2005-12-06 Xylb  xylulokinase homolog (H. influenzae)  Xylb_predicted  xylulokinase homolog (H. influenzae) (predicted)  Symbol and Name updated 1559027 APPROVED
2005-01-12 Xylb_predicted  xylulokinase homolog (H. influenzae) (predicted)      Symbol and Name status set to approved 70820 APPROVED