Nek9 (NIMA-related kinase 9) - Rat Genome Database

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Gene: Nek9 (NIMA-related kinase 9) Rattus norvegicus
Analyze
Symbol: Nek9
Name: NIMA-related kinase 9
RGD ID: 1307187
Description: Predicted to enable protein kinase activator activity; protein kinase binding activity; and protein serine/threonine kinase activity. Predicted to be involved in mitotic cell cycle. Predicted to be located in exocytic vesicle. Predicted to colocalize with centrosome. Human ortholog(s) of this gene implicated in lethal congenital contracture syndrome. Orthologous to human NEK9 (NIMA related kinase 9); INTERACTS WITH 17alpha-ethynylestradiol; 2,4-dinitrotoluene; 2,6-dinitrotoluene.
Type: protein-coding
RefSeq Status: VALIDATED
Previously known as: LOC299204; NIMA (never in mitosis gene a)- related kinase 9; serine/threonine-protein kinase Nek9
RGD Orthologs
Human
Mouse
Chinchilla
Bonobo
Dog
Squirrel
Pig
Green Monkey
Naked Mole-Rat
Alliance Genes
More Info more info ...
Latest Assembly: mRatBN7.2 - mRatBN7.2 Assembly
Position:
Rat AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
mRatBN7.26104,944,056 - 104,984,538 (-)NCBImRatBN7.2mRatBN7.2
mRatBN7.2 Ensembl6104,944,056 - 104,984,538 (-)EnsemblmRatBN7.2 Ensembl
UTH_Rnor_SHR_Utx6105,113,660 - 105,154,136 (-)NCBIRnor_SHR
UTH_Rnor_SHRSP_BbbUtx_1.06105,412,532 - 105,453,008 (-)NCBIRnor_SHRSP
UTH_Rnor_WKY_Bbb_1.06104,782,190 - 104,822,670 (-)NCBIRnor_WKY
Rnor_6.06109,121,524 - 109,162,433 (-)NCBIRnor6.0Rnor_6.0rn6Rnor6.0
Rnor_6.0 Ensembl6109,124,330 - 109,162,267 (-)EnsemblRnor6.0rn6Rnor6.0
Rnor_5.06116,390,523 - 116,435,517 (+)NCBIRnor5.0Rnor_5.0rn5Rnor5.0
RGSC_v3.46109,345,437 - 109,389,174 (-)NCBIRGSC3.4RGSC_v3.4rn4RGSC3.4
RGSC_v3.16109,351,642 - 109,394,468 (-)NCBI
Celera6102,766,488 - 102,807,366 (-)NCBICelera
Cytogenetic Map6q31NCBI
JBrowse: View Region in Genome Browser (JBrowse)
Model


Gene Ontology Annotations     Click to see Annotation Detail View

Biological Process

Cellular Component
centrosome  (IBA,ISO)
exocytic vesicle  (ISO)

Molecular Function

References

References - curated
# Reference Title Reference Citation
1. Phylogenetic-based propagation of functional annotations within the Gene Ontology consortium. Gaudet P, etal., Brief Bioinform. 2011 Sep;12(5):449-62. doi: 10.1093/bib/bbr042. Epub 2011 Aug 27.
2. GOA pipeline RGD automated data pipeline
3. ClinVar Automated Import and Annotation Pipeline RGD automated import pipeline for ClinVar variants, variant-to-disease annotations and gene-to-disease annotations
4. Data Import for Chemical-Gene Interactions RGD automated import pipeline for gene-chemical interactions
Additional References at PubMed
PMID:18504258   PMID:19001501   PMID:20873783  


Genomics

Comparative Map Data
Nek9
(Rattus norvegicus - Norway rat)
Rat AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
mRatBN7.26104,944,056 - 104,984,538 (-)NCBImRatBN7.2mRatBN7.2
mRatBN7.2 Ensembl6104,944,056 - 104,984,538 (-)EnsemblmRatBN7.2 Ensembl
UTH_Rnor_SHR_Utx6105,113,660 - 105,154,136 (-)NCBIRnor_SHR
UTH_Rnor_SHRSP_BbbUtx_1.06105,412,532 - 105,453,008 (-)NCBIRnor_SHRSP
UTH_Rnor_WKY_Bbb_1.06104,782,190 - 104,822,670 (-)NCBIRnor_WKY
Rnor_6.06109,121,524 - 109,162,433 (-)NCBIRnor6.0Rnor_6.0rn6Rnor6.0
Rnor_6.0 Ensembl6109,124,330 - 109,162,267 (-)EnsemblRnor6.0rn6Rnor6.0
Rnor_5.06116,390,523 - 116,435,517 (+)NCBIRnor5.0Rnor_5.0rn5Rnor5.0
RGSC_v3.46109,345,437 - 109,389,174 (-)NCBIRGSC3.4RGSC_v3.4rn4RGSC3.4
RGSC_v3.16109,351,642 - 109,394,468 (-)NCBI
Celera6102,766,488 - 102,807,366 (-)NCBICelera
Cytogenetic Map6q31NCBI
NEK9
(Homo sapiens - human)
Human AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
GRCh381475,079,353 - 75,127,202 (-)NCBIGRCh38GRCh38hg38GRCh38
GRCh38.p13 Ensembl1475,079,353 - 75,127,344 (-)EnsemblGRCh38hg38GRCh38
GRCh371475,546,056 - 75,593,751 (-)NCBIGRCh37GRCh37hg19GRCh37
Build 361474,620,728 - 74,663,389 (-)NCBINCBI36Build 36hg18NCBI36
Build 341474,620,727 - 74,663,389NCBI
Celera1455,586,461 - 55,631,402 (-)NCBICelera
Cytogenetic Map14q24.3NCBI
HuRef1455,716,982 - 55,761,929 (-)NCBIHuRef
CHM1_11475,486,591 - 75,531,543 (-)NCBICHM1_1
T2T-CHM13v2.01469,288,752 - 69,336,432 (-)NCBIT2T-CHM13v2.0
Nek9
(Mus musculus - house mouse)
Mouse AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
GRCm391285,346,288 - 85,386,136 (-)NCBIGRCm39GRCm39mm39
GRCm39 Ensembl1285,346,288 - 85,386,136 (-)EnsemblGRCm39 Ensembl
GRCm381285,299,514 - 85,339,362 (-)NCBIGRCm38GRCm38mm10GRCm38
GRCm38.p6 Ensembl1285,299,514 - 85,339,362 (-)EnsemblGRCm38mm10GRCm38
MGSCv371286,640,464 - 86,680,312 (-)NCBIGRCm37MGSCv37mm9NCBIm37
MGSCv361286,188,616 - 86,228,440 (-)NCBIMGSCv36mm8
Celera1286,757,015 - 86,796,871 (-)NCBICelera
Cytogenetic Map12D2NCBI
Nek9
(Chinchilla lanigera - long-tailed chinchilla)
Chinchilla AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
ChiLan1.0 EnsemblNW_004955523446,748 - 482,246 (+)EnsemblChiLan1.0
ChiLan1.0NW_004955523443,832 - 483,397 (+)NCBIChiLan1.0ChiLan1.0
NEK9
(Pan paniscus - bonobo/pygmy chimpanzee)
Bonobo AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
PanPan1.11474,478,387 - 74,522,320 (-)NCBIpanpan1.1PanPan1.1panPan2
PanPan1.1 Ensembl1474,478,387 - 74,522,320 (-)Ensemblpanpan1.1panPan2
Mhudiblu_PPA_v01455,633,667 - 55,678,670 (-)NCBIMhudiblu_PPA_v0Mhudiblu_PPA_v0panPan3
NEK9
(Canis lupus familiaris - dog)
Dog AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
CanFam3.1848,180,203 - 48,218,430 (-)NCBICanFam3.1CanFam3.1canFam3CanFam3.1
CanFam3.1 Ensembl848,183,071 - 48,257,646 (-)EnsemblCanFam3.1canFam3CanFam3.1
Dog10K_Boxer_Tasha847,864,320 - 47,902,676 (-)NCBIDog10K_Boxer_Tasha
ROS_Cfam_1.0848,407,827 - 48,446,234 (-)NCBIROS_Cfam_1.0
ROS_Cfam_1.0 Ensembl848,407,827 - 48,446,233 (-)EnsemblROS_Cfam_1.0 Ensembl
UMICH_Zoey_3.1848,078,603 - 48,116,941 (-)NCBIUMICH_Zoey_3.1
UNSW_CanFamBas_1.0848,099,872 - 48,138,217 (-)NCBIUNSW_CanFamBas_1.0
UU_Cfam_GSD_1.0848,498,571 - 48,536,910 (-)NCBIUU_Cfam_GSD_1.0
Nek9
(Ictidomys tridecemlineatus - thirteen-lined ground squirrel)
Squirrel AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
HiC_Itri_2NW_02440864027,807,701 - 27,847,583 (+)NCBIHiC_Itri_2
SpeTri2.0 EnsemblNW_0049364884,220,429 - 4,261,432 (-)EnsemblSpeTri2.0
SpeTri2.0NW_0049364884,221,099 - 4,261,401 (-)NCBISpeTri2.0SpeTri2.0SpeTri2.0
NEK9
(Sus scrofa - pig)
Pig AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
Sscrofa11.1 Ensembl798,275,979 - 98,322,055 (-)EnsemblSscrofa11.1susScr11Sscrofa11.1
Sscrofa11.1798,275,973 - 98,322,072 (-)NCBISscrofa11.1Sscrofa11.1susScr11Sscrofa11.1
Sscrofa10.27104,120,094 - 104,166,188 (-)NCBISscrofa10.2Sscrofa10.2susScr3
NEK9
(Chlorocebus sabaeus - green monkey)
Green Monkey AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
ChlSab1.12452,297,211 - 52,342,359 (-)NCBIChlSab1.1ChlSab1.1chlSab2
ChlSab1.1 Ensembl2452,296,261 - 52,342,724 (-)EnsemblChlSab1.1ChlSab1.1 EnsemblchlSab2
Vero_WHO_p1.0NW_02366605340,527,142 - 40,572,456 (-)NCBIVero_WHO_p1.0Vero_WHO_p1.0
Nek9
(Heterocephalus glaber - naked mole-rat)
Naked Mole-rat AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
HetGla 1.0NW_00462473426,557,208 - 26,600,777 (+)NCBIHetGla_female_1.0HetGla 1.0hetGla2

Variants

.
Variants in Nek9
317 total Variants
miRNA Target Status

Predicted Target Of
Summary Value
Count of predictions:549
Count of miRNA genes:251
Interacting mature miRNAs:341
Transcripts:ENSRNOT00000068017
Prediction methods:Microtar, Miranda, Targetscan
Result types:miRGate_prediction

The detailed report is available here: Full Report CSV TAB Printer

miRNA Target Status data imported from miRGate (http://mirgate.bioinfo.cnio.es/).
For more information about miRGate, see PMID:25858286 or access the full paper here.


QTLs in Region (mRatBN7.2)
The following QTLs overlap with this region.    Full Report CSV TAB Printer Gviewer
RGD IDSymbolNameLODP ValueTraitSub TraitChrStartStopSpecies
4145119Mcs25Mammary carcinoma susceptibility QTL 250.0001mammary gland integrity trait (VT:0010552)ratio of deaths to total study population during a period of time (CMO:0001023)610894415110548006Rat
1576309Emca7Estrogen-induced mammary cancer QTL 74mammary gland integrity trait (VT:0010552)mammary tumor number (CMO:0000343)615107216107351382Rat
12801471Schws9Schwannoma susceptibility QTL 9nervous system integrity trait (VT:0010566)percentage of study population developing trigeminal nerve neurilemmomas during a period of time (CMO:0002017)661747639106747639Rat
731173Uae22Urinary albumin excretion QTL 2210.1urine albumin amount (VT:0002871)urine albumin excretion rate (CMO:0000757)665531555140994061Rat
2290393Uae37Urinary albumin excretion QTL 370.0001urine albumin amount (VT:0002871)urine albumin excretion rate (CMO:0000757)665531555140994061Rat
1641904Alcrsp4Alcohol response QTL 4response to alcohol trait (VT:0010489)duration of loss of righting reflex (CMO:0002289)667262953112262953Rat
1300075Glom7Glomerulus QTL 75.60.0000002kidney glomerulus morphology trait (VT:0005325)count of superficial glomeruli not directly contacting the kidney surface (CMO:0001002)671201409116201409Rat
1331789Rf37Renal function QTL 373.224kidney blood vessel physiology trait (VT:0100012)absolute change in renal vascular resistance (CMO:0001900)672202632115200186Rat
70196BpQTLcluster7Blood pressure QTL cluster 76.82arterial blood pressure trait (VT:2000000)systolic blood pressure (CMO:0000004)672202632117202632Rat
724536Uae7Urinary albumin excretion QTL 73.5urine albumin amount (VT:0002871)urine albumin level (CMO:0000130)672202632130729475Rat
1331799Bp211Blood pressure QTL 2113.66407arterial blood pressure trait (VT:2000000)mean arterial blood pressure (CMO:0000009)672202632130919985Rat
1581550Pur8Proteinuria QTL 8urine total protein amount (VT:0000032)urine total protein excretion rate (CMO:0000756)672227641130729205Rat
1581563Uae33Urinary albumin excretion QTL 33urine albumin amount (VT:0002871)urine albumin excretion rate (CMO:0000757)672227641130729205Rat
724524Uae2Urinary albumin excretion QTL 22.70.0005urine albumin amount (VT:0002871)urine albumin excretion rate (CMO:0000757)673463459109394713Rat
737827Hcar11Hepatocarcinoma resistance QTL 114.4liver integrity trait (VT:0010547)liver tumorous lesion number (CMO:0001068)682523650110548006Rat
1576302Schws4Schwannoma susceptibility QTL 40.0078nervous system integrity trait (VT:0010566)percentage of study population developing trigeminal nerve neurilemmomas during a period of time (CMO:0002017)683190345106747639Rat
738034Anxrr5Anxiety related response QTL 55.9exploratory behavior trait (VT:0010471)percentage of entries into a discrete space in an experimental apparatus (CMO:0000961)684130881129130881Rat
10054138Gmadr3Adrenal mass QTL 33.680.00045adrenal gland mass (VT:0010420)both adrenal glands wet weight (CMO:0000164)685140138130140138Rat
10054123Srcrt6Stress Responsive Cort QTL 62.50.0043blood corticosterone amount (VT:0005345)plasma corticosterone level (CMO:0001173)685140138130140138Rat
724513Uae14Urinary albumin excretion QTL 146.5urine albumin amount (VT:0002871)urine albumin excretion rate (CMO:0000757)685311061133478515Rat
1300076Glom8Glomerulus QTL 870.000000009kidney glomerulus morphology trait (VT:0005325)count of superficial glomeruli directly contacting the kidney surface (CMO:0001001)686894788131894788Rat
2303624Vencon5Ventilatory control QTL 54.45respiration trait (VT:0001943)minute ventilation (CMO:0000132)688047916133047916Rat
1331722Thshl1Thyroid stimulating hormone level QTL 111.70.0001blood thyroid-stimulating hormone amount (VT:0005119)serum thyroid stimulating hormone level (CMO:0001248)689762877106752806Rat
1331725Bp212Blood pressure QTL 2123.52475arterial blood pressure trait (VT:2000000)mean arterial blood pressure (CMO:0000009)693701310128713626Rat
61414Pia3Pristane induced arthritis QTL 34.5joint integrity trait (VT:0010548)post-insult time to onset of experimental arthritis (CMO:0001450)694968928137848904Rat
12801411Schws8Schwannoma susceptibility QTL 8nervous system integrity trait (VT:0010566)percentage of study population developing trigeminal nerve neurilemmomas during a period of time (CMO:0002017)694968928139968928Rat
8552796Vie3Viral induced encephalitis QTL 32.6brain integrity trait (VT:0010579)encephalitis incidence/prevalence measurement (CMO:0002361)696833997140994061Rat
1358355Srcrt4Stress Responsive Cort QTL 46.39blood corticosterone amount (VT:0005345)plasma corticosterone level (CMO:0001173)6100364669140994061Rat
2313399Anxrr28Anxiety related response QTL 28aggression-related behavior trait (VT:0015014)tameness/aggressiveness composite score (CMO:0002136)6100671796132340886Rat
1331797Bp213Blood pressure QTL 2133.291arterial blood pressure trait (VT:2000000)mean arterial blood pressure (CMO:0000009)6104085867128713626Rat

Markers in Region
RH142370  
Rat AssemblyChrPosition (strand)SourceJBrowse
mRatBN7.26104,944,305 - 104,944,485 (+)MAPPERmRatBN7.2
Rnor_6.06109,122,209 - 109,122,388NCBIRnor6.0
Rnor_5.06116,434,653 - 116,434,832UniSTSRnor5.0
RGSC_v3.46109,346,122 - 109,346,301UniSTSRGSC3.4
Celera6102,767,173 - 102,767,352UniSTS
RH 3.4 Map6742.39UniSTS
Cytogenetic Map6q31UniSTS
BE104575  
Rat AssemblyChrPosition (strand)SourceJBrowse
mRatBN7.26104,944,895 - 104,945,074 (+)MAPPERmRatBN7.2
Rnor_6.06109,122,799 - 109,122,977NCBIRnor6.0
Rnor_5.06116,434,064 - 116,434,242UniSTSRnor5.0
RGSC_v3.46109,346,712 - 109,346,890UniSTSRGSC3.4
Celera6102,767,763 - 102,767,941UniSTS
RH 3.4 Map6742.4UniSTS
Cytogenetic Map6q31UniSTS
RH134586  
Rat AssemblyChrPosition (strand)SourceJBrowse
mRatBN7.26104,945,599 - 104,945,808 (+)MAPPERmRatBN7.2
Rnor_6.06109,123,503 - 109,123,711NCBIRnor6.0
Rnor_5.06116,433,330 - 116,433,538UniSTSRnor5.0
RGSC_v3.46109,347,416 - 109,347,624UniSTSRGSC3.4
Celera6102,768,467 - 102,768,675UniSTS
RH 3.4 Map6732.39UniSTS
Cytogenetic Map6q31UniSTS


Expression


RNA-SEQ Expression
High: > 1000 TPM value   Medium: Between 11 and 1000 TPM
Low: Between 0.5 and 10 TPM   Below Cutoff: < 0.5 TPM

alimentary part of gastrointestinal system circulatory system endocrine system exocrine system hemolymphoid system hepatobiliary system integumental system musculoskeletal system nervous system renal system reproductive system respiratory system appendage
High
Medium 3 42 52 36 19 36 8 11 74 35 40 11 8
Low 1 5 5 5 1
Below cutoff

Sequence


Reference Sequences
RefSeq Acc Id: ENSRNOT00000077518   ⟹   ENSRNOP00000074679
RefSeq Status:
Type: CODING
Position:
Rat AssemblyChrPosition (strand)Source
mRatBN7.2 Ensembl6104,947,898 - 104,984,538 (-)Ensembl
Rnor_6.0 Ensembl6109,124,330 - 109,162,267 (-)Ensembl
RefSeq Acc Id: ENSRNOT00000106509   ⟹   ENSRNOP00000096294
RefSeq Status:
Type: CODING
Position:
Rat AssemblyChrPosition (strand)Source
mRatBN7.2 Ensembl6104,944,056 - 104,984,538 (-)Ensembl
RefSeq Acc Id: NM_001106747   ⟹   NP_001100217
RefSeq Status: VALIDATED
Type: CODING
Position:
Rat AssemblyChrPosition (strand)Source
mRatBN7.26104,944,056 - 104,984,538 (-)NCBI
Rnor_6.06109,121,524 - 109,162,409 (-)NCBI
Rnor_5.06116,390,523 - 116,435,517 (+)NCBI
RGSC_v3.46109,345,437 - 109,389,174 (-)RGD
Celera6102,766,488 - 102,807,366 (-)RGD
Sequence:
RefSeq Acc Id: XM_039112049   ⟹   XP_038967977
RefSeq Status:
Type: CODING
Position:
Rat AssemblyChrPosition (strand)Source
mRatBN7.26104,944,056 - 104,984,536 (-)NCBI
Protein Sequences
Protein RefSeqs NP_001100217 (Get FASTA)   NCBI Sequence Viewer  
  XP_038967977 (Get FASTA)   NCBI Sequence Viewer  
GenBank Protein EDL81568 (Get FASTA)   NCBI Sequence Viewer  
Reference Sequences
RefSeq Acc Id: NP_001100217   ⟸   NM_001106747
- UniProtKB: A0A8I6AQL7 (UniProtKB/TrEMBL)
- Sequence:
RefSeq Acc Id: ENSRNOP00000074679   ⟸   ENSRNOT00000077518
RefSeq Acc Id: XP_038967977   ⟸   XM_039112049
- Peptide Label: isoform X1
RefSeq Acc Id: ENSRNOP00000096294   ⟸   ENSRNOT00000106509
Protein Domains
Protein kinase

Protein Structures
Name Modeler Protein Id AA Range Protein Structure
AF-A0A0G2K8N9-F1-model_v2 AlphaFold A0A0G2K8N9 1-958 view protein structure

Transcriptome

eQTL   View at Phenogen
WGCNA   View at Phenogen
Tissue/Strain Expression   View at Phenogen


Additional Information

Database Acc Id Source(s)
AGR Gene RGD:1307187 AgrOrtholog
BioCyc Gene G2FUF-36788 BioCyc
Ensembl Genes ENSRNOG00000058572 Ensembl, ENTREZGENE, UniProtKB/TrEMBL
Ensembl Protein ENSRNOP00000074679.2 UniProtKB/TrEMBL
  ENSRNOP00000096294 ENTREZGENE
  ENSRNOP00000096294.1 UniProtKB/TrEMBL
Ensembl Transcript ENSRNOT00000077518.2 UniProtKB/TrEMBL
  ENSRNOT00000106509 ENTREZGENE
  ENSRNOT00000106509.1 UniProtKB/TrEMBL
Gene3D-CATH 2.130.10.30 UniProtKB/TrEMBL
InterPro Kinase-like_dom UniProtKB/TrEMBL
  Nek9_STKc UniProtKB/TrEMBL
  Prot_kinase_cat_dom UniProtKB/TrEMBL
  Reg_chr_condens UniProtKB/TrEMBL
  Reg_csome_cond/b-lactamase_inh UniProtKB/TrEMBL
  Ser/Thr_prot_kinase_AS UniProtKB/TrEMBL
NCBI Gene 299204 ENTREZGENE
Pfam Pkinase UniProtKB/TrEMBL
  RCC1 UniProtKB/TrEMBL
PhenoGen Nek9 PhenoGen
PROSITE PROTEIN_KINASE_DOM UniProtKB/TrEMBL
  PROTEIN_KINASE_ST UniProtKB/TrEMBL
  RCC1_3 UniProtKB/TrEMBL
SMART S_TKc UniProtKB/TrEMBL
Superfamily-SCOP Kinase_like UniProtKB/TrEMBL
  RCC1/BLIP-II UniProtKB/TrEMBL
UniProt A0A0G2K8N9_RAT UniProtKB/TrEMBL
  A0A8I6AQL7 ENTREZGENE, UniProtKB/TrEMBL


Nomenclature History
Date Current Symbol Current Name Previous Symbol Previous Name Description Reference Status
2012-11-21 Nek9  NIMA-related kinase 9  Nek9  NIMA (never in mitosis gene a)- related kinase 9  Nomenclature updated to reflect human and mouse nomenclature 1299863 APPROVED
2008-04-30 Nek9  NIMA (never in mitosis gene a)- related kinase 9   Nek9_predicted  NIMA (never in mitosis gene a)- related kinase 9 (predicted)  'predicted' is removed 2292626 APPROVED
2005-01-12 Nek9_predicted  NIMA (never in mitosis gene a)- related kinase 9 (predicted)      Symbol and Name status set to approved 70820 APPROVED