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The Vertebrate Trait Ontology (VT) is being developed as a collaborative effort between the Rat Genome Database at the Medical College of Wisconsin (RGD,, Mouse Genome Informatics at the Jackson Laboratory ( and the Animal QTL database at Iowa State University ( For more information about this vocabulary, or to request additions or changes, please contact us (

Term:surface structure morphology trait
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Accession:VT:0015066 term browser browse the term
Definition:Any measurable or observable characteristic related to the shape, structure, color or pattern of the organism subdivision which is the collection of anatomical structures on the body surface.

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coat/hair pigmentation trait term browser
Symbol Object Name Evidence Notes Source PubMed Reference(s) RGD Reference(s) Position
G Asip agouti signaling protein ISO Coat colour, agouti OMIA PMID:12692166 PMID:15520882 PMID:15965787 PMID:17630272 PMID:21321476 PMID:21846741 PMID:22433982 PMID:22524303 PMID:23519866 PMID:32635139 NCBI chr 3:150,492,010...150,579,870
Ensembl chr 3:150,574,473...150,579,870
JBrowse link
G Defb14 defensin beta 14 ISO Coat colour, dominant black OMIA PMID:12692166 PMID:15649147 PMID:17483404 PMID:17947548 PMID:19197024 PMID:21321476 PMID:24098367 NCBI chr16:75,647,523...75,650,913
Ensembl chr16:75,647,680...75,648,538
JBrowse link
G Mc1r melanocortin 1 receptor ISO Coat colour, melanistic mask OMIA PMID:12692165 PMID:21353269 PMID:29932470 PMID:33672409 NCBI chr19:56,215,420...56,219,987 JBrowse link
G Pmel premelanosome protein ISO Coat colour, merle OMIA PMID:330141 PMID:340189 PMID:418699 PMID:3099512 PMID:3998446 PMID:4031467 PMID:6323572 PMID:6410614 PMID:6817463 PMID:7006993 PMID:12580795 PMID:16407134 PMID:16504149 PMID:17186850 PMID:19192156 PMID:27387721 PMID:30071510 PMID:30123327 PMID:30235206 PMID:31238727 PMID:32560567 NCBI chr 7:3,133,347...3,143,827
Ensembl chr 7:3,133,506...3,143,911
JBrowse link
G Psmb7 proteasome 20S subunit beta 7 ISO Coat colour, harlequin OMIA PMID:3270445 PMID:3998446 PMID:18513894 PMID:21256207 NCBI chr 3:22,890,072...22,951,718
Ensembl chr 3:22,871,105...22,952,063
JBrowse link
G Raly RALY heterogeneous nuclear ribonucleoprotein ISO Coat colour, saddle tan vs black-and-tan OMIA PMID:23519866 PMID:33672409 NCBI chr 3:150,322,834...150,387,345
Ensembl chr 3:150,323,116...150,387,332
JBrowse link

Term paths to the root
Path 1
Term Annotations click to browse term
  vertebrate trait 2385
    organism subdivision trait 186
      surface structure trait 186
        surface structure morphology trait 186
          appendage morphology trait + 142
          beak morphology trait 0
          cloaca morphology trait 0
          ear morphology trait + 0
          eye morphology trait + 14
          integumentary system morphology trait + 30
          lip morphology trait + 0
          nose morphology trait + 0
          opercular flap morphology trait 0
          snout morphology trait + 0
paths to the root