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ONTOLOGY REPORT - ANNOTATIONS


Term:nuclear DNA-directed RNA polymerase complex
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Accession:GO:0055029 term browser browse the term
Definition:A protein complex, located in the nucleus, that possesses DNA-directed RNA polymerase activity.


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alpha DNA polymerase:primase complex term browser
Symbol Object Name Qualifiers Evidence Notes Source PubMed Reference(s) RGD Reference(s) Position
G Pola1 DNA polymerase alpha 1, catalytic subunit IDA
IBA
ISO
ISS
IEA
IC
PMID:21873635
(MGI:1203906|PMID:9584195)
(PMID:2175912), (PMID:22465957)
GO_REF:0000024
GO_REF:0000107
RGD
UniProt
Ensembl
PMID:2175912 PMID:9584195 PMID:22465957, PMID:7670930, PMID:21873635, PMID:7503737 GO_REF:0000024 GO_REF:0000107 MGI:1203906, RGD:1580605, RGD:13792537, RGD:2307013 NCBI chr  X:62,382,604...62,698,830
Ensembl chr  X:62,382,608...62,698,830
JBrowse link
G Pola2 DNA polymerase alpha 2, accessory subunit TAS
IBA
ISO
IEA
IC
PMID:21873635
(MGI:1203906|PMID:9584195)
(PMID:22465957)
GO_REF:0000107
RGD
Ensembl
PMID:9584195 PMID:22465957, PMID:11536367, PMID:21873635, PMID:7503737 GO_REF:0000107 MGI:1203906, RGD:633584, RGD:13792537, RGD:2307013 NCBI chr 1:221,297,417...221,321,601
Ensembl chr 1:221,273,538...221,321,570
JBrowse link
G Prim1 DNA primase subunit 1 IBA PMID:21873635 GO_Central PMID:21873635 RGD:13792537 NCBI chr 7:2,435,524...2,451,169
Ensembl chr 7:2,435,553...2,451,165
JBrowse link
G Prim2 DNA primase subunit 2 IBA PMID:21873635 GO_Central PMID:21873635 RGD:13792537 NCBI chr 9:38,535,435...38,745,058
Ensembl chr 9:38,536,920...38,745,099
JBrowse link
Cdc73/Paf1 complex term browser
Symbol Object Name Qualifiers Evidence Notes Source PubMed Reference(s) RGD Reference(s) Position
G Cdc73 cell division cycle 73 ISS
IEA
IBA
ISO
GO_REF:0000024
GO_REF:0000107
PMID:21873635
(PMID:16307923), (PMID:18987311), (PMID:20178742)
UniProt
Ensembl
RGD
PMID:16307923 PMID:18987311 PMID:20178742, PMID:21873635 GO_REF:0000024 GO_REF:0000107, RGD:13792537 NCBI chr13:60,403,771...60,496,511
Ensembl chr13:60,403,771...60,496,511
JBrowse link
G Ctr9 CTR9 homolog, Paf1/RNA polymerase II complex component colocalizes_with ISO
IBA
(PMID:16307923), (PMID:20178742)
UniProtKB:Q8JZM7 (MGI:6203683|PMID:27749823)
PMID:21873635
RGD PMID:16307923 PMID:20178742 PMID:27749823, PMID:21873635 MGI:6203683, RGD:13792537 NCBI chr 1:175,839,140...175,869,167
Ensembl chr 1:175,839,108...175,869,165
JBrowse link
G Leo1 LEO1 homolog, Paf1/RNA polymerase II complex component ISS
ISO
GO_REF:0000024
(PMID:16307923), (PMID:18987311), (PMID:20178742)
UniProt
RGD
PMID:16307923 PMID:18987311 PMID:20178742 GO_REF:0000024 NCBI chr 8:82,380,906...82,405,293
Ensembl chr 8:82,380,757...82,405,644
JBrowse link
G Paf1 PAF1 homolog, Paf1/RNA polymerase II complex component ISS
ISO
IBA
GO_REF:0000024
UniProtKB:Q8JZM7 (MGI:6203683|PMID:27749823)
PMID:21873635
(PMID:16307923), (PMID:20178742)
UniProt
RGD
PMID:16307923 PMID:20178742 PMID:27749823, PMID:21873635 GO_REF:0000024 MGI:6203683, RGD:13792537 NCBI chr 1:85,362,421...85,367,999
Ensembl chr 1:85,362,268...85,368,031
JBrowse link
G Pex2 peroxisomal biogenesis factor 2 ISO (PMID:18987311) RGD PMID:18987311 NCBI chr 2:98,251,756...98,269,185
Ensembl chr 2:98,252,925...98,269,184
JBrowse link
G Rtf1 Rtf1, Paf1/RNA polymerase II complex component, homolog (S. cerevisiae) ISO
IBA
(PMID:20178742)
UniProtKB:Q8JZM7 (MGI:6203683|PMID:27749823)
PMID:21873635
RGD PMID:20178742 PMID:27749823, PMID:21873635 MGI:6203683, RGD:13792537 NCBI chr 3:111,478,350...111,537,941
Ensembl chr 3:111,478,382...111,537,941
JBrowse link
G Setd5 SET domain containing 5 colocalizes_with ISO (MGI:5812827|PMID:27864380) RGD PMID:27864380 MGI:5812827 NCBI chr 4:145,017,549...145,095,247
Ensembl chr 4:145,017,608...145,095,245
JBrowse link
G Wdr61 WD repeat domain 61 ISO
ISS
(PMID:20178742)
UniProtKB:Q8JZM7 (MGI:6203683|PMID:27749823)
GO_REF:0000024
RGD
UniProt
PMID:20178742 PMID:27749823 GO_REF:0000024 MGI:6203683 NCBI chr 8:59,296,339...59,313,663
Ensembl chr 8:59,296,346...59,313,574
JBrowse link
core TFIIH complex portion of holo TFIIH complex term browser
Symbol Object Name Qualifiers Evidence Notes Source PubMed Reference(s) RGD Reference(s) Position
G Gtf2h2 general transcription factor IIH subunit 2 ISS GO_REF:0000024 UniProt GO_REF:0000024 NCBI chr 2:30,494,718...30,522,716
Ensembl chr 2:30,494,825...30,522,727
JBrowse link
G Gtf2h3 general transcription factor IIH subunit 3 ISO
ISS
(PMID:27193682)
GO_REF:0000024
RGD
UniProt
PMID:27193682 GO_REF:0000024 NCBI chr12:37,427,556...37,444,027
Ensembl chr12:37,427,546...37,444,282
JBrowse link
G Gtf2h4 general transcription factor 2H subunit 4 ISO (PMID:27193682) RGD PMID:27193682 NCBI chr20:3,582,688...3,588,373
Ensembl chr20:3,582,742...3,588,375
JBrowse link
RNA polymerase I complex term browser
Symbol Object Name Qualifiers Evidence Notes Source PubMed Reference(s) RGD Reference(s) Position
G AABR07006025.1 IBA PMID:21873635 GO_Central PMID:21873635 RGD:13792537 Ensembl chr 1:214,015,219...214,024,153 JBrowse link
G AABR07042789.1 IBA PMID:21873635 GO_Central PMID:21873635 RGD:13792537 Ensembl chr19:8,943,753...8,943,956 JBrowse link
G AC109542.1 IBA PMID:21873635 GO_Central PMID:21873635 RGD:13792537 Ensembl chr 1:214,452,127...214,455,039 JBrowse link
G LOC100912534 DNA-directed RNA polymerases I, II, and III subunit RPABC3-like IBA PMID:21873635 GO_Central PMID:21873635 RGD:13792537 NCBI chr11:82,856,537...82,860,991
Ensembl chr11:82,856,529...82,860,977
JBrowse link
G Polr1a RNA polymerase I subunit A IDA
IBA
ISO
IEA
PMID:21873635
(MGI:3047492|PMID:15226435), (MGI:894558|PMID:9236775)
GO_REF:0000107
RGD
Ensembl
PMID:9236775 PMID:15226435, PMID:9422795, PMID:21873635 GO_REF:0000107 MGI:3047492 MGI:894558, RGD:2312782, RGD:13792537 NCBI chr 4:99,822,964...99,903,969
Ensembl chr 4:99,823,252...99,887,132
JBrowse link
G Polr1b RNA polymerase I subunit B IDA
IBA
ISO
IEA
PMID:21873635
(MGI:894558|PMID:9236775)
GO_REF:0000107
RGD
Ensembl
PMID:9236775, PMID:9422795, PMID:21873635 GO_REF:0000107 MGI:894558, RGD:2312782, RGD:13792537 NCBI chr 3:121,632,043...121,656,506
Ensembl chr 3:121,632,043...121,656,506
JBrowse link
G Polr1c RNA polymerase I and III subunit C ISO
IBA
(MGI:69752|PMID:7929437)
PMID:21873635
RGD PMID:7929437, PMID:21873635 MGI:69752, RGD:13792537 NCBI chr 9:17,120,759...17,124,871
Ensembl chr 9:17,120,759...17,124,871
JBrowse link
G Polr1d RNA polymerase I and III subunit D ISO (MGI:3047492|PMID:15226435) RGD PMID:15226435 MGI:3047492 NCBI chr12:9,693,828...9,727,416
Ensembl chr12:9,693,779...9,727,489
JBrowse link
G Polr1e RNA polymerase I subunit E ISO
IBA
(MGI:1928312|PMID:8641287)
PMID:21873635
RGD PMID:8641287, PMID:21873635 MGI:1928312, RGD:13792537 NCBI chr 5:60,572,573...60,588,097
Ensembl chr 5:60,572,577...60,588,240
JBrowse link
G Polr1f RNA polymerase I subunit F IBA PMID:21873635 GO_Central PMID:21873635 RGD:13792537 NCBI chr 6:52,751,106...52,768,574
Ensembl chr 6:52,751,106...52,768,574
JBrowse link
G Polr1g RNA polymerase I subunit G ISO
IBA
(MGI:3047492|PMID:15226435)
PMID:21873635
RGD PMID:15226435, PMID:21873635 MGI:3047492, RGD:13792537 NCBI chr 1:80,267,725...80,271,001
Ensembl chr 1:80,267,725...80,271,001
JBrowse link
G Polr1h RNA polymerase I subunit H IBA PMID:21873635 GO_Central PMID:21873635 RGD:13792537 NCBI chr20:2,088,518...2,092,436
Ensembl chr20:2,088,533...2,092,436
JBrowse link
G Polr2e RNA polymerase II, I and III subunit E IBA PMID:21873635 GO_Central PMID:21873635 RGD:13792537 NCBI chr 7:12,532,785...12,536,797
Ensembl chr 7:12,532,895...12,536,791
JBrowse link
G Polr2f RNA polymerase II, I and III subunit F TAS
IBA
PMID:21873635 RGD
GO_Central
PMID:10393248, PMID:21873635 RGD:1299534, RGD:13792537 NCBI chr 7:120,380,543...120,392,214
Ensembl chr 7:120,380,544...120,390,954
JBrowse link
G Polr2h RNA polymerase II, I and III subunit H IBA PMID:21873635 GO_Central PMID:21873635 RGD:13792537 NCBI chr11:83,877,760...83,882,208
Ensembl chr11:83,878,006...83,882,117
JBrowse link
G Polr2k RNA polymerase II, I and III subunit K IBA PMID:21873635 GO_Central PMID:21873635 RGD:13792537 NCBI chr 7:74,939,010...74,992,594
Ensembl chr 7:74,989,223...74,992,582
JBrowse link
G Polr2l-ps1 polymerase (RNA) II (DNA directed) polypeptide L, pseudogene 1 IBA PMID:21873635 GO_Central PMID:21873635 RGD:13792537 NCBI chr19:8,943,599...8,943,966 JBrowse link
RNA polymerase II, core complex term browser
Symbol Object Name Qualifiers Evidence Notes Source PubMed Reference(s) RGD Reference(s) Position
G AABR07006025.1 IBA PMID:21873635 GO_Central PMID:21873635 RGD:13792537 Ensembl chr 1:214,015,219...214,024,153 JBrowse link
G AABR07042789.1 IBA PMID:21873635 GO_Central PMID:21873635 RGD:13792537 Ensembl chr19:8,943,753...8,943,956 JBrowse link
G AC109542.1 IBA PMID:21873635 GO_Central PMID:21873635 RGD:13792537 Ensembl chr 1:214,452,127...214,455,039 JBrowse link
G LOC100912534 DNA-directed RNA polymerases I, II, and III subunit RPABC3-like IBA PMID:21873635 GO_Central PMID:21873635 RGD:13792537 NCBI chr11:82,856,537...82,860,991
Ensembl chr11:82,856,529...82,860,977
JBrowse link
G LOC103690054 DNA-directed RNA polymerase II subunit RPB9 IBA PMID:21873635 GO_Central PMID:21873635 RGD:13792537 NCBI chr 1:91,041,173...91,042,258
Ensembl chr 1:91,041,214...91,042,230
Ensembl chr 1:91,041,214...91,042,230
JBrowse link
G Polr2a RNA polymerase II subunit A ISO
IBA
(PMID:9852112)
PMID:21873635
RGD PMID:9852112, PMID:21873635 RGD:13792537 NCBI chr10:56,339,277...56,364,888
Ensembl chr10:56,339,284...56,365,084
JBrowse link
G Polr2b RNA polymerase II subunit B ISO
IBA
(PMID:9852112)
PMID:21873635
RGD PMID:9852112, PMID:21873635 RGD:13792537 NCBI chr14:33,070,104...33,107,792
Ensembl chr14:33,070,078...33,107,776
JBrowse link
G Polr2c RNA polymerase II subunit C ISO
IBA
(PMID:9852112)
PMID:21873635
RGD PMID:9852112, PMID:21873635 RGD:13792537 NCBI chr19:10,575,060...10,581,842
Ensembl chr19:10,575,060...10,581,842
JBrowse link
G Polr2d RNA polymerase II subunit D ISO
IBA
(PMID:9528765), (PMID:9852112)
PMID:21873635
RGD PMID:9528765 PMID:9852112, PMID:21873635 RGD:13792537 NCBI chr18:24,570,762...24,577,894
Ensembl chr18:24,570,762...24,577,894
JBrowse link
G Polr2e RNA polymerase II, I and III subunit E ISO
IBA
ISS
(PMID:9852112)
PMID:21873635
GO_REF:0000024
RGD
UniProt
PMID:9852112, PMID:21873635 GO_REF:0000024, RGD:13792537 NCBI chr 7:12,532,785...12,536,797
Ensembl chr 7:12,532,895...12,536,791
JBrowse link
G Polr2f RNA polymerase II, I and III subunit F TAS
IBA
ISS
ISO
PMID:21873635
GO_REF:0000024
(PMID:9852112)
UniProt
RGD
PMID:9852112, PMID:10393248, PMID:21873635 GO_REF:0000024, RGD:1299534, RGD:13792537 NCBI chr 7:120,380,543...120,392,214
Ensembl chr 7:120,380,544...120,390,954
JBrowse link
G Polr2g RNA polymerase II subunit G ISO
IBA
ISS
(PMID:9852112)
PMID:21873635
GO_REF:0000024
RGD
UniProt
PMID:9852112, PMID:21873635 GO_REF:0000024, RGD:13792537 NCBI chr 1:224,991,250...224,998,355
Ensembl chr 1:224,991,222...224,994,653
JBrowse link
G Polr2h RNA polymerase II, I and III subunit H ISO
IBA
(PMID:9852112)
PMID:21873635
RGD PMID:9852112, PMID:21873635 RGD:13792537 NCBI chr11:83,877,760...83,882,208
Ensembl chr11:83,878,006...83,882,117
JBrowse link
G Polr2i RNA polymerase II subunit I ISO
IBA
(PMID:9852112)
PMID:21873635
RGD PMID:9852112, PMID:21873635 RGD:13792537 NCBI chr 1:88,686,812...88,688,249
Ensembl chr 1:88,686,731...88,688,258
JBrowse link
G Polr2j RNA polymerase II subunit J ISO
IBA
(PMID:9852112)
PMID:21873635
(MGI:86690|PMID:9099876)
RGD PMID:9099876 PMID:9852112, PMID:21873635 MGI:86690, RGD:13792537 NCBI chr12:23,587,222...23,592,774
Ensembl chr12:23,587,222...23,592,774
JBrowse link
G Polr2k RNA polymerase II, I and III subunit K ISO
IBA
(PMID:9852112)
PMID:21873635
RGD PMID:9852112, PMID:21873635 RGD:13792537 NCBI chr 7:74,939,010...74,992,594
Ensembl chr 7:74,989,223...74,992,582
JBrowse link
G Polr2l RNA polymerase II, I and III subunit L ISO (PMID:9852112) RGD PMID:9852112 NCBI chr 1:214,451,966...214,453,974 JBrowse link
G Polr2l-ps1 polymerase (RNA) II (DNA directed) polypeptide L, pseudogene 1 IBA PMID:21873635 GO_Central PMID:21873635 RGD:13792537 NCBI chr19:8,943,599...8,943,966 JBrowse link
G Polr2m RNA polymerase II subunit M ISO UniProtKB:P19387|UniProtKB:P24928|UniProtKB:P30876 (PMID:30190596) RGD PMID:30190596 NCBI chr 8:77,985,190...77,992,621
Ensembl chr 8:77,985,190...78,096,390
JBrowse link
RNA polymerase II, holoenzyme term browser
Symbol Object Name Qualifiers Evidence Notes Source PubMed Reference(s) RGD Reference(s) Position
G AABR07043557.1 IBA PMID:21873635 GO_Central PMID:21873635 RGD:13792537 Ensembl chr19:29,320,701...29,324,203 JBrowse link
G Myo6 myosin VI ISO (PMID:16949370) RGD PMID:16949370 NCBI chr 8:87,583,649...87,731,271
Ensembl chr 8:87,630,916...87,731,269
JBrowse link
G Myzap myocardial zonula adherens protein IEA GO_REF:0000002 InterPro GO_REF:0000002 NCBI chr 8:78,009,019...78,096,409
Ensembl chr 8:77,985,190...78,096,390
JBrowse link
G Polr2m RNA polymerase II subunit M IEA
IBA
ISS
ISO
GO_REF:0000002
PMID:21873635
GO_REF:0000024
(PMID:22231121)
InterPro
UniProt
RGD
PMID:22231121, PMID:21873635 GO_REF:0000002 GO_REF:0000024, RGD:13792537 NCBI chr 8:77,985,190...77,992,621
Ensembl chr 8:77,985,190...78,096,390
JBrowse link
G Recql5 RecQ like helicase 5 ISO (PMID:22231121) RGD PMID:22231121 NCBI chr10:104,443,735...104,482,889
Ensembl chr10:104,443,736...104,482,838
JBrowse link
G Rpap2 RNA polymerase II associated protein 2 ISS
ISO
GO_REF:0000024
(PMID:22137580), (PMID:22231121)
UniProt
RGD
PMID:22137580 PMID:22231121 GO_REF:0000024 NCBI chr14:3,125,595...3,204,590
Ensembl chr14:3,124,622...3,204,588
JBrowse link
G Rprd1a regulation of nuclear pre-mRNA domain containing 1A ISO
IBA
(PMID:22231121)
PMID:21873635
RGD PMID:22231121, PMID:21873635 RGD:13792537 NCBI chr18:16,450,160...16,497,913
Ensembl chr18:16,453,369...16,497,886
JBrowse link
G Rprd1b regulation of nuclear pre-mRNA domain containing 1B ISO
IBA
(PMID:22231121)
PMID:21873635
RGD PMID:22231121, PMID:21873635 RGD:13792537 NCBI chr 3:154,507,029...154,553,312
Ensembl chr 3:154,507,035...154,553,312
JBrowse link
G Rprd2 regulation of nuclear pre-mRNA domain containing 2 ISO
IBA
(PMID:22231121)
PMID:21873635
RGD PMID:22231121, PMID:21873635 RGD:13792537 NCBI chr 2:197,882,666...197,937,844
Ensembl chr 2:197,886,160...197,935,567
JBrowse link
RNA polymerase III complex term browser
Symbol Object Name Qualifiers Evidence Notes Source PubMed Reference(s) RGD Reference(s) Position
G AABR07006025.1 IBA PMID:21873635 GO_Central PMID:21873635 RGD:13792537 Ensembl chr 1:214,015,219...214,024,153 JBrowse link
G AABR07042789.1 IBA PMID:21873635 GO_Central PMID:21873635 RGD:13792537 Ensembl chr19:8,943,753...8,943,956 JBrowse link
G AC109542.1 IBA PMID:21873635 GO_Central PMID:21873635 RGD:13792537 Ensembl chr 1:214,452,127...214,455,039 JBrowse link
G Crcp CGRP receptor component IEA
IBA
GO_REF:0000002
PMID:21873635
InterPro PMID:21873635 GO_REF:0000002, RGD:13792537 NCBI chr12:30,125,237...30,160,857
Ensembl chr12:30,125,168...30,180,115
JBrowse link
G LOC100912534 DNA-directed RNA polymerases I, II, and III subunit RPABC3-like IBA PMID:21873635 GO_Central PMID:21873635 RGD:13792537 NCBI chr11:82,856,537...82,860,991
Ensembl chr11:82,856,529...82,860,977
JBrowse link
G Polr1c RNA polymerase I and III subunit C ISO
IBA
(PMID:24107381)
PMID:21873635
RGD PMID:24107381, PMID:21873635 RGD:13792537 NCBI chr 9:17,120,759...17,124,871
Ensembl chr 9:17,120,759...17,124,871
JBrowse link
G Polr2e RNA polymerase II, I and III subunit E IBA PMID:21873635 GO_Central PMID:21873635 RGD:13792537 NCBI chr 7:12,532,785...12,536,797
Ensembl chr 7:12,532,895...12,536,791
JBrowse link
G Polr2f RNA polymerase II, I and III subunit F TAS
IBA
PMID:21873635 RGD
GO_Central
PMID:10393248, PMID:21873635 RGD:1299534, RGD:13792537 NCBI chr 7:120,380,543...120,392,214
Ensembl chr 7:120,380,544...120,390,954
JBrowse link
G Polr2h RNA polymerase II, I and III subunit H IBA PMID:21873635 GO_Central PMID:21873635 RGD:13792537 NCBI chr11:83,877,760...83,882,208
Ensembl chr11:83,878,006...83,882,117
JBrowse link
G Polr2k RNA polymerase II, I and III subunit K IBA PMID:21873635 GO_Central PMID:21873635 RGD:13792537 NCBI chr 7:74,939,010...74,992,594
Ensembl chr 7:74,989,223...74,992,582
JBrowse link
G Polr2l-ps1 polymerase (RNA) II (DNA directed) polypeptide L, pseudogene 1 IBA PMID:21873635 GO_Central PMID:21873635 RGD:13792537 NCBI chr19:8,943,599...8,943,966 JBrowse link
G Polr3a RNA polymerase III subunit A ISO
IBA
(PMID:24107381)
PMID:21873635
RGD PMID:24107381, PMID:21873635 RGD:13792537 NCBI chr16:717,821...756,002
Ensembl chr16:716,683...755,990
JBrowse link
G Polr3b RNA polymerase III subunit B ISO
IBA
(PMID:24107381)
PMID:21873635
RGD PMID:24107381, PMID:21873635 RGD:13792537 NCBI chr 7:24,745,051...24,851,227
Ensembl chr 7:24,745,051...24,851,227
JBrowse link
G Polr3c RNA polymerase III subunit C ISO
IBA
(PMID:24107381)
PMID:21873635
RGD PMID:24107381, PMID:21873635 RGD:13792537 NCBI chr 2:198,836,282...198,852,368
Ensembl chr 2:198,836,285...198,852,161
JBrowse link
G Polr3d RNA polymerase III subunit D ISO
IBA
(PMID:24107381)
PMID:21873635
RGD PMID:24107381, PMID:21873635 RGD:13792537 NCBI chr15:52,129,471...52,134,140
Ensembl chr15:52,129,471...52,134,140
JBrowse link
G Polr3e RNA polymerase III subunit E ISO
IBA
(PMID:24107381)
PMID:21873635
RGD PMID:24107381, PMID:21873635 RGD:13792537 NCBI chr 1:190,852,954...190,881,539
Ensembl chr 1:190,852,985...190,881,108
JBrowse link
G Polr3f RNA polymerase III subunit F ISO
IBA
(PMID:24107381)
PMID:21873635
RGD PMID:24107381, PMID:21873635 RGD:13792537 NCBI chr 3:138,684,685...138,701,055
Ensembl chr 3:138,684,685...138,701,055
JBrowse link
G Polr3g RNA polymerase III subunit G ISO
IBA
(PMID:24107381)
PMID:21873635
RGD PMID:24107381, PMID:21873635 RGD:13792537 NCBI chr 2:9,537,642...9,578,551
Ensembl chr 2:9,537,642...9,578,310
JBrowse link
G Polr3gl RNA polymerase III subunit GL ISO
IBA
(PMID:24107381)
PMID:21873635
RGD PMID:24107381, PMID:21873635 RGD:13792537 NCBI chr 2:198,698,044...198,719,609
Ensembl chr 2:198,702,587...198,719,202
JBrowse link
G Polr3h RNA polymerase III subunit H ISO
IBA
(PMID:12391170)
PMID:21873635
RGD PMID:12391170, PMID:21873635 RGD:13792537 NCBI chr 7:123,146,248...123,156,558
Ensembl chr 7:123,146,249...123,156,558
JBrowse link
G Polr3k RNA polymerase III subunit K IBA PMID:21873635 GO_Central PMID:21873635 RGD:13792537 NCBI chr 3:177,374,812...177,379,009
Ensembl chr 3:177,374,812...177,379,008
JBrowse link
transcription factor TFIIA complex term browser
Symbol Object Name Qualifiers Evidence Notes Source PubMed Reference(s) RGD Reference(s) Position
G Gtf2a1 general transcription factor 2A subunit 1 ISO
IBA
(PMID:7724559)
PMID:21873635
(MGI:3583836|PMID:15927180)
RGD PMID:7724559 PMID:15927180, PMID:21873635 MGI:3583836, RGD:13792537 NCBI chr 6:115,321,110...115,352,659
Ensembl chr 6:115,320,467...115,352,681
JBrowse link
G Gtf2a1l general transcription factor 2A subunit 1 like ISO
IBA
(MGI:3583836|PMID:15927180)
PMID:21873635
RGD PMID:15927180, PMID:21873635 MGI:3583836, RGD:13792537 NCBI chr 6:12,412,198...12,468,613
Ensembl chr 6:12,415,805...12,468,596
JBrowse link
G Gtf2a2 general transcription factor 2A subunit 2 IEA
IBA
ISO
GO_REF:0000002
PMID:21873635
(PMID:7724559), (PMID:7958899), (PMID:7958900)
InterPro
RGD
PMID:7724559 PMID:7958899 PMID:7958900, PMID:21873635 GO_REF:0000002, RGD:13792537 NCBI chr 8:76,422,341...76,435,587
Ensembl chr 8:76,422,359...76,435,473
JBrowse link
G LOC103695118 transcription initiation factor IIA subunit 2 IBA
IEA
PMID:21873635
GO_REF:0000002
InterPro PMID:21873635 GO_REF:0000002, RGD:13792537 Ensembl chr 8:76,422,359...76,435,473 JBrowse link
G Tbp TATA box binding protein ISO (PMID:7724559) RGD PMID:7724559 NCBI chr 1:57,491,381...57,509,335
Ensembl chr 1:57,491,643...57,508,449
JBrowse link
G Tbpl1 TATA-box binding protein like 1 ISO (MGI:3583836|PMID:15927180) RGD PMID:15927180 MGI:3583836 NCBI chr 1:23,977,154...24,002,512
Ensembl chr 1:23,977,688...24,002,506
JBrowse link
transcription factor TFIID complex term browser
Symbol Object Name Qualifiers Evidence Notes Source PubMed Reference(s) RGD Reference(s) Position
G AABR07063323.1 IBA PMID:21873635 GO_Central PMID:21873635 RGD:13792537 Ensembl chr 6:28,557,633...28,557,848 JBrowse link
G Ercc2 ERCC excision repair 2, TFIIH core complex helicase subunit ISO (PMID:27193682) RGD PMID:27193682 NCBI chr 1:80,293,574...80,307,334
Ensembl chr 1:80,293,566...80,307,344
JBrowse link
G Ercc3 ERCC excision repair 3, TFIIH core complex helicase subunit ISO (PMID:27193682) RGD PMID:27193682 NCBI chr18:25,037,668...25,068,380
Ensembl chr18:25,037,625...25,068,389
JBrowse link
G Gtf2a1 general transcription factor 2A subunit 1 ISO (PMID:27193682) RGD PMID:27193682 NCBI chr 6:115,321,110...115,352,659
Ensembl chr 6:115,320,467...115,352,681
JBrowse link
G Gtf2a2 general transcription factor 2A subunit 2 ISO (PMID:27193682) RGD PMID:27193682 NCBI chr 8:76,422,341...76,435,587
Ensembl chr 8:76,422,359...76,435,473
JBrowse link
G Gtf2b general transcription factor IIB IEA
ISO
GO_REF:0000107
(PMID:27193682)
Ensembl
RGD
PMID:27193682 GO_REF:0000107 NCBI chr 2:248,709,433...248,727,741
Ensembl chr 2:248,715,281...248,727,741
JBrowse link
G Gtf2e1 general transcription factor IIE subunit 1 ISO (PMID:27193682) RGD PMID:27193682 NCBI chr11:66,078,786...66,111,394
Ensembl chr11:66,078,820...66,110,848
JBrowse link
G Gtf2e2 general transcription factor IIE subunit 2 ISO (PMID:27193682) RGD PMID:27193682 NCBI chr16:62,113,846...62,164,339
Ensembl chr16:62,113,854...62,164,135
JBrowse link
G Gtf2f1 general transcription factor IIF subunit 1 ISO (PMID:27193682) RGD PMID:27193682 NCBI chr 9:10,032,806...10,042,392
Ensembl chr 9:10,033,011...10,042,418
JBrowse link
G Gtf2h3 general transcription factor IIH subunit 3 ISO (PMID:27193682) RGD PMID:27193682 NCBI chr12:37,427,556...37,444,027
Ensembl chr12:37,427,546...37,444,282
JBrowse link
G Gtf2h4 general transcription factor 2H subunit 4 ISO (PMID:27193682) RGD PMID:27193682 NCBI chr20:3,582,688...3,588,373
Ensembl chr20:3,582,742...3,588,375
JBrowse link
G Gtf2h5 general transcription factor IIH subunit 5 ISO
IBA
(PMID:27193682)
PMID:21873635
RGD PMID:27193682, PMID:21873635 RGD:13792537 NCBI chr 1:46,978,345...46,985,032
Ensembl chr 1:46,978,458...46,985,032
JBrowse link
G Taf1 TATA-box binding protein associated factor 1 ISO (PMID:14580349), (PMID:24927529), (PMID:25412659), (PMID:27007846), (PMID:29111974), (PMID:8625415) RGD PMID:8625415 PMID:14580349 PMID:24927529 PMID:25412659 PMID:27007846 PMID:29111974 NCBI chr  X:71,412,291...71,486,456
Ensembl chr  X:71,412,289...71,485,085
JBrowse link
G Taf10 TATA-box binding protein associated factor 10 ISO (PMID:14580349), (PMID:15099517), (PMID:29111974), (PMID:7729427), (PMID:7923369) RGD PMID:7729427 PMID:7923369 PMID:14580349 PMID:15099517 PMID:29111974 NCBI chr 1:170,585,160...170,586,428
Ensembl chr 1:170,585,167...170,586,603
JBrowse link
G Taf11 TATA-box binding protein associated factor 11 IBA
ISO
PMID:21873635
(PMID:14580349), (PMID:29111974), (PMID:7729427), (PMID:8670810)
RGD PMID:7729427 PMID:8670810 PMID:14580349 PMID:29111974, PMID:21873635 RGD:13792537 NCBI chr20:7,476,648...7,484,417
Ensembl chr20:7,476,651...7,482,747
JBrowse link
G Taf12 TATA-box binding protein associated factor 12 IBA
ISO
PMID:21873635
(PMID:14580349), (PMID:29111974)
RGD PMID:14580349 PMID:29111974, PMID:21873635 RGD:13792537 NCBI chr 5:150,459,701...150,476,521
Ensembl chr 5:150,459,713...150,475,186
JBrowse link
G Taf13 TATA-box binding protein associated factor 13 ISO
IBA
(PMID:29111974), (PMID:7729427)
PMID:21873635
RGD PMID:7729427 PMID:29111974, PMID:21873635 RGD:13792537 NCBI chr 2:211,359,623...211,370,285
Ensembl chr 2:211,359,623...211,370,284
JBrowse link
G Taf2 TATA-box binding protein associated factor 2 IBA
ISO
PMID:21873635
(PMID:10409744), (PMID:14580349), (PMID:27007846), (PMID:29111974), (PMID:9774672)
RGD PMID:9774672 PMID:10409744 PMID:14580349 PMID:27007846 PMID:29111974, PMID:21873635 RGD:13792537 NCBI chr 7:94,698,987...94,755,924
Ensembl chr 7:94,698,987...94,755,924
JBrowse link
G Taf3 TATA-box binding protein associated factor 3 ISO (PMID:11438666), (PMID:29111974) RGD PMID:11438666 PMID:29111974 NCBI chr17:72,240,785...72,393,448
Ensembl chr17:72,209,373...72,394,271
JBrowse link
G Taf4 TATA-box binding protein associated factor 4 ISO
IBA
(PMID:14580349), (PMID:29111974)
PMID:21873635
RGD PMID:14580349 PMID:29111974, PMID:21873635 RGD:13792537 NCBI chr 3:175,300,842...175,364,655
Ensembl chr 3:175,301,879...175,327,689
JBrowse link
G Taf4b TATA-box binding protein associated factor 4b ISO
IBA
(PMID:29111974)
PMID:21873635
RGD PMID:29111974, PMID:21873635 RGD:13792537 NCBI chr18:6,206,939...6,338,794
Ensembl chr18:6,207,280...6,318,219
JBrowse link
G Taf5 TATA-box binding protein associated factor 5 IBA
ISO
PMID:21873635
(PMID:14580349), (PMID:29111974), (PMID:9045704)
RGD PMID:9045704 PMID:14580349 PMID:29111974, PMID:21873635 RGD:13792537 NCBI chr 1:266,844,480...266,859,965
Ensembl chr 1:266,844,480...266,859,965
JBrowse link
G Taf6 TATA-box binding protein associated factor 6 IEA
IBA
ISO
GO_REF:0000107
PMID:21873635
(PMID:14580349), (PMID:27007846), (PMID:29111974), (PMID:8262073)
Ensembl
RGD
PMID:8262073 PMID:14580349 PMID:27007846 PMID:29111974, PMID:21873635 GO_REF:0000107, RGD:13792537 NCBI chr12:19,320,269...19,328,706
Ensembl chr12:19,320,271...19,328,637
JBrowse link
G Taf6l TATA-box binding protein associated factor 6 like IEA GO_REF:0000002 InterPro GO_REF:0000002 NCBI chr 1:224,974,238...224,986,819
Ensembl chr 1:224,974,208...224,986,291
JBrowse link
G Taf7 TATA-box binding protein associated factor 7 ISO
IBA
(PMID:14580349), (PMID:24927529), (PMID:25412659), (PMID:27007846), (PMID:29111974), (PMID:7824954)
PMID:21873635
(MGI:1342782|PMID:10438527)
RGD PMID:7824954 PMID:10438527 PMID:14580349 PMID:24927529 PMID:25412659 PMID:27007846 PMID:29111974, PMID:21873635 MGI:1342782, RGD:13792537 NCBI chr18:30,779,149...30,781,297
Ensembl chr18:30,779,150...30,781,292
JBrowse link
G Taf7l TATA-box binding protein associated factor 7-like IBA PMID:21873635 GO_Central PMID:21873635 RGD:13792537 NCBI chr  X:105,292,338...105,308,554
Ensembl chr  X:105,292,338...105,308,554
JBrowse link
G Taf7l-ps1 TAF7-like RNA polymerase II, TATA box binding protein (TBP)-associated factor, pseudogene 1 IBA PMID:21873635 GO_Central PMID:21873635 RGD:13792537 NCBI chr 7:59,138,768...59,139,910 JBrowse link
G Taf7l2 Taf7l2 IBA PMID:21873635 GO_Central PMID:21873635 RGD:13792537 Ensembl chr 7:59,138,768...59,139,910 JBrowse link
G Taf8 TATA-box binding protein associated factor 8 ISO
IBA
(PMID:14580349), (PMID:27007846), (PMID:29111974)
PMID:21873635
RGD PMID:14580349 PMID:27007846 PMID:29111974, PMID:21873635 RGD:13792537 NCBI chr 9:15,513,048...15,532,992
Ensembl chr 9:15,513,063...15,529,800
JBrowse link
G Taf9 TATA-box binding protein associated factor 9 IBA
ISO
PMID:21873635
(PMID:14580349), (PMID:15899866), (PMID:29111974)
RGD PMID:14580349 PMID:15899866 PMID:29111974, PMID:21873635 RGD:13792537 NCBI chr 2:30,664,407...30,668,414
Ensembl chr 2:30,664,639...30,668,414
JBrowse link
G Taf9b TATA-box binding protein associated factor 9b IBA
ISO
PMID:21873635
(PMID:15899866), (PMID:29111974)
RGD PMID:15899866 PMID:29111974, PMID:21873635 RGD:13792537 NCBI chr  X:77,281,234...77,295,238
Ensembl chr  X:77,281,210...77,295,426
JBrowse link
G Tbp TATA box binding protein IDA
ISO
(PMID:14580349), (PMID:27007846), (PMID:27193682), (PMID:29111974), (PMID:7729427) RGD PMID:7729427 PMID:14580349 PMID:27007846 PMID:27193682 PMID:29111974, PMID:7980586 RGD:2306552 NCBI chr 1:57,491,381...57,509,335
Ensembl chr 1:57,491,643...57,508,449
JBrowse link
G Tcea1 transcription elongation factor A1 ISO (PMID:27193682) RGD PMID:27193682 NCBI chr 5:14,511,183...14,548,494
Ensembl chr 5:14,511,183...14,548,429
JBrowse link
G Tp53 tumor protein p53 colocalizes_with ISO (PMID:15053879) RGD PMID:15053879 NCBI chr10:56,186,299...56,198,449
Ensembl chr10:56,187,020...56,198,449
JBrowse link
transcription factor TFIIE complex term browser
Symbol Object Name Qualifiers Evidence Notes Source PubMed Reference(s) RGD Reference(s) Position
G Gtf2e1 general transcription factor IIE subunit 1 IBA PMID:21873635 GO_Central PMID:21873635 RGD:13792537 NCBI chr11:66,078,786...66,111,394
Ensembl chr11:66,078,820...66,110,848
JBrowse link
G Gtf2e2 general transcription factor IIE subunit 2 IBA PMID:21873635 GO_Central PMID:21873635 RGD:13792537 NCBI chr16:62,113,846...62,164,339
Ensembl chr16:62,113,854...62,164,135
JBrowse link
transcription factor TFIIF complex term browser
Symbol Object Name Qualifiers Evidence Notes Source PubMed Reference(s) RGD Reference(s) Position
G Gtf2f1 general transcription factor IIF subunit 1 IBA PMID:21873635 GO_Central PMID:21873635 RGD:13792537 NCBI chr 9:10,032,806...10,042,392
Ensembl chr 9:10,033,011...10,042,418
JBrowse link
G Gtf2f2 general transcription factor IIF subunit 2 IEA
IBA
GO_REF:0000002
PMID:21873635
InterPro PMID:21873635 GO_REF:0000002, RGD:13792537 NCBI chr15:57,942,057...58,068,892
Ensembl chr15:57,942,059...58,068,892
JBrowse link
transcription factor TFIIH holo complex term browser
Symbol Object Name Qualifiers Evidence Notes Source PubMed Reference(s) RGD Reference(s) Position
G AABR07063323.1 IBA PMID:21873635 GO_Central PMID:21873635 RGD:13792537 Ensembl chr 6:28,557,633...28,557,848 JBrowse link
G Ccnh cyclin H ISO
IBA
ISS
(PMID:9852112)
PMID:21873635
GO_REF:0000024
RGD
UniProt
PMID:9852112, PMID:21873635 GO_REF:0000024, RGD:13792537 NCBI chr 2:13,593,100...13,613,910
Ensembl chr 2:13,593,100...13,613,910
JBrowse link
G Cdk7 cyclin-dependent kinase 7 IDA
ISS
ISO
GO_REF:0000024
(PMID:9852112)
UniProt
RGD
PMID:9852112, PMID:7553866 GO_REF:0000024, RGD:9590319 NCBI chr 2:30,710,642...30,735,522
Ensembl chr 2:30,710,643...30,735,514
JBrowse link
G Ercc2 ERCC excision repair 2, TFIIH core complex helicase subunit ISO (PMID:9852112) RGD PMID:9852112 NCBI chr 1:80,293,574...80,307,334
Ensembl chr 1:80,293,566...80,307,344
JBrowse link
G Ercc3 ERCC excision repair 3, TFIIH core complex helicase subunit ISS
ISO
IBA
GO_REF:0000024
(PMID:9852112)
PMID:21873635
UniProt
RGD
PMID:9852112, PMID:21873635 GO_REF:0000024, RGD:13792537 NCBI chr18:25,037,668...25,068,380
Ensembl chr18:25,037,625...25,068,389
JBrowse link
G Gtf2f2 general transcription factor IIF subunit 2 IDA RGD PMID:7553866 RGD:9590319 NCBI chr15:57,942,057...58,068,892
Ensembl chr15:57,942,059...58,068,892
JBrowse link
G Gtf2h1 general transcription factor IIH subunit 1 IDA
IBA
ISO
PMID:21873635
(PMID:9852112)
RGD PMID:9852112, PMID:7553866, PMID:21873635 RGD:9590319, RGD:13792537 NCBI chr 1:102,849,618...102,877,939
Ensembl chr 1:102,849,889...102,877,944
JBrowse link
G Gtf2h2 general transcription factor IIH subunit 2 ISS
IBA
GO_REF:0000024
PMID:21873635
UniProt PMID:21873635 GO_REF:0000024, RGD:13792537 NCBI chr 2:30,494,718...30,522,716
Ensembl chr 2:30,494,825...30,522,727
JBrowse link
G Gtf2h3 general transcription factor IIH subunit 3 ISO
IBA
ISS
(PMID:9852112)
PMID:21873635
GO_REF:0000024
RGD
UniProt
PMID:9852112, PMID:21873635 GO_REF:0000024, RGD:13792537 NCBI chr12:37,427,556...37,444,027
Ensembl chr12:37,427,546...37,444,282
JBrowse link
G Gtf2h4 general transcription factor 2H subunit 4 ISO
IBA
(PMID:9852112)
PMID:21873635
RGD PMID:9852112, PMID:21873635 RGD:13792537 NCBI chr20:3,582,688...3,588,373
Ensembl chr20:3,582,742...3,588,375
JBrowse link
G Gtf2h5 general transcription factor IIH subunit 5 IBA PMID:21873635 GO_Central PMID:21873635 RGD:13792537 NCBI chr 1:46,978,345...46,985,032
Ensembl chr 1:46,978,458...46,985,032
JBrowse link
G Mnat1 MNAT1 component of CDK activating kinase ISO
IBA
(PMID:9852112)
PMID:21873635
RGD PMID:9852112, PMID:21873635 RGD:13792537 NCBI chr 6:95,995,341...96,153,331
Ensembl chr 6:96,007,805...96,153,330
JBrowse link
transcription factor TFIIK complex term browser
Symbol Object Name Qualifiers Evidence Notes Source PubMed Reference(s) RGD Reference(s) Position
G Ccnh cyclin H IBA
ISO
PMID:21873635
(PMID:8692841)
RGD PMID:8692841, PMID:21873635 RGD:13792537 NCBI chr 2:13,593,100...13,613,910
Ensembl chr 2:13,593,100...13,613,910
JBrowse link
G Cdk7 cyclin-dependent kinase 7 IBA
IEA
ISO
PMID:21873635
GO_REF:0000107
(PMID:8692841)
Ensembl
RGD
PMID:8692841, PMID:21873635 GO_REF:0000107, RGD:13792537 NCBI chr 2:30,710,642...30,735,522
Ensembl chr 2:30,710,643...30,735,514
JBrowse link
G Mnat1 MNAT1 component of CDK activating kinase ISO (PMID:8692841) RGD PMID:8692841 NCBI chr 6:95,995,341...96,153,331
Ensembl chr 6:96,007,805...96,153,330
JBrowse link
transcription factor TFTC complex term browser
Symbol Object Name Qualifiers Evidence Notes Source PubMed Reference(s) RGD Reference(s) Position
G Kat2a lysine acetyltransferase 2A ISO (PMID:10373431) RGD PMID:10373431 NCBI chr10:88,611,586...88,619,558
Ensembl chr10:88,611,586...88,619,537
JBrowse link
G LOC690283 similar to TAF5-like RNA polymerase II, p300/CBP-associated factor (PCAF)-associated factor IEA GO_REF:0000002 InterPro GO_REF:0000002 NCBI chr 1:60,105,788...60,108,103
Ensembl chr 1:60,106,034...60,107,039
JBrowse link
G Supt3h SPT3 homolog, SAGA and STAGA complex component ISO (PMID:10373431) RGD PMID:10373431 NCBI chr 9:18,249,565...18,604,814
Ensembl chr 9:18,319,246...18,371,856
JBrowse link
G Tada3 transcriptional adaptor 3 ISO (PMID:10373431) RGD PMID:10373431 NCBI chr 4:145,318,417...145,330,291
Ensembl chr 4:145,318,435...145,329,878
JBrowse link
G Taf10 TATA-box binding protein associated factor 10 ISO (PMID:10373431), (PMID:9603525) RGD PMID:9603525 PMID:10373431 NCBI chr 1:170,585,160...170,586,428
Ensembl chr 1:170,585,167...170,586,603
JBrowse link
G Taf12 TATA-box binding protein associated factor 12 ISO (PMID:9603525) RGD PMID:9603525 NCBI chr 5:150,459,701...150,476,521
Ensembl chr 5:150,459,713...150,475,186
JBrowse link
G Taf2 TATA-box binding protein associated factor 2 ISO (PMID:10373431) RGD PMID:10373431 NCBI chr 7:94,698,987...94,755,924
Ensembl chr 7:94,698,987...94,755,924
JBrowse link
G Taf4 TATA-box binding protein associated factor 4 ISO (PMID:10373431), (PMID:9603525) RGD PMID:9603525 PMID:10373431 NCBI chr 3:175,300,842...175,364,655
Ensembl chr 3:175,301,879...175,327,689
JBrowse link
G Taf5 TATA-box binding protein associated factor 5 ISO (PMID:10373431), (PMID:9603525) RGD PMID:9603525 PMID:10373431 NCBI chr 1:266,844,480...266,859,965
Ensembl chr 1:266,844,480...266,859,965
JBrowse link
G Taf5l TATA-box binding protein associated factor 5 like ISO (PMID:10373431) RGD PMID:10373431 NCBI chr19:56,781,854...56,801,005
Ensembl chr19:56,781,859...56,799,445
JBrowse link
G Taf6 TATA-box binding protein associated factor 6 IEA
ISO
GO_REF:0000107
(PMID:9603525)
Ensembl
RGD
PMID:9603525 GO_REF:0000107 NCBI chr12:19,320,269...19,328,706
Ensembl chr12:19,320,271...19,328,637
JBrowse link
G Taf7 TATA-box binding protein associated factor 7 ISO (PMID:9603525) RGD PMID:9603525 NCBI chr18:30,779,149...30,781,297
Ensembl chr18:30,779,150...30,781,292
JBrowse link
G Taf9 TATA-box binding protein associated factor 9 ISO
IBA
(PMID:15899866)
PMID:21873635
RGD PMID:15899866, PMID:21873635 RGD:13792537 NCBI chr 2:30,664,407...30,668,414
Ensembl chr 2:30,664,639...30,668,414
JBrowse link
G Taf9b TATA-box binding protein associated factor 9b ISO (PMID:15899866) RGD PMID:15899866 NCBI chr  X:77,281,234...77,295,238
Ensembl chr  X:77,281,210...77,295,426
JBrowse link
G Trrap transformation/transcription domain-associated protein ISO (PMID:10373431) RGD PMID:10373431 NCBI chr12:11,537,950...11,627,563
Ensembl chr12:11,537,950...11,627,563
JBrowse link

Term paths to the root
Path 1
Term Annotations click to browse term
  cellular_component 20615
    protein-containing complex 6107
      nuclear protein-containing complex 1237
        nuclear DNA-directed RNA polymerase complex 107
          RNA polymerase I complex 17
          RNA polymerase II, core complex 19
          RNA polymerase II, holoenzyme + 84
          RNA polymerase III complex 21
          RNA polymerase IV complex 0
          RNA polymerase V complex 0
          alpha DNA polymerase:primase complex 4
Path 2
Term Annotations click to browse term
  cellular_component 20615
    protein-containing complex 6107
      catalytic complex 1497
        transferase complex 822
          transferase complex, transferring phosphorus-containing groups 266
            RNA polymerase complex 110
              DNA-directed RNA polymerase complex 108
                nuclear DNA-directed RNA polymerase complex 107
                  RNA polymerase I complex 17
                  RNA polymerase II, core complex 19
                  RNA polymerase II, holoenzyme + 84
                  RNA polymerase III complex 21
                  RNA polymerase IV complex 0
                  RNA polymerase V complex 0
                  alpha DNA polymerase:primase complex 4
paths to the root