ONTOLOGY REPORT - ANNOTATIONS
Term: proteasome complex
Accession: GO:0000502
browse the term
Definition: A large multisubunit complex which catalyzes protein degradation, found in eukaryotes, archaea and some bacteria. In eukaryotes, this complex consists of the barrel shaped proteasome core complex and one or two associated proteins or complexes that act in regulating entry into or exit from the core.
Synonyms: exact_synonym: proteasome
narrow_synonym: 26S proteasome
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Adrm1
ADRM1 26S proteasome ubiquitin receptor
part_of
ISO IEA ISS
(PMID:16990800), (PMID:17323924) GO_REF:0000107 GO_REF:0000043 GO_REF:0000024
RGD Ensembl UniProt
PMID:16990800 PMID:17323924
GO_REF:0000024 GO_REF:0000043 GO_REF:0000107
NCBI chr 3:167,265,435...167,270,197
Ensembl chr 3:167,265,296...167,270,195
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Dnajb2
DnaJ heat shock protein family (Hsp40) member B2
located_in
ISO
(PMID:15936278)
RGD
PMID:15936278
NCBI chr 9:76,731,060...76,739,278
Ensembl chr 9:76,731,065...76,739,277
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Hspb1
heat shock protein family B (small) member 1
IDA
RGD
PMID:12897149
RGD:1304417
NCBI chr12:20,794,014...20,795,675
Ensembl chr12:20,794,028...20,795,743
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Prickle1
prickle planar cell polarity protein 1
part_of
IEA ISO
GO_REF:0000107 (PMID:25190059)
Ensembl RGD
PMID:25190059
GO_REF:0000107
NCBI chr 7:124,639,142...124,735,027
Ensembl chr 7:124,639,142...124,658,113
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Psma1
proteasome 20S subunit alpha 1
part_of
ISO
(PMID:17323924) (PMID:16973439)
RGD
PMID:16973439 PMID:17323924
NCBI chr 1:168,442,421...168,453,440
Ensembl chr 1:168,440,936...168,453,472
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Psma2
proteasome 20S subunit alpha 2
part_of
ISO
(PMID:17323924)
RGD
PMID:17323924
NCBI chr17:50,554,536...50,564,923
Ensembl chr17:50,551,924...50,564,938
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Psma3
proteasome 20S subunit alpha 3
part_of
ISO
(PMID:17323924), (PMID:26524591)
RGD
PMID:17323924 PMID:26524591
NCBI chr 6:89,483,741...89,503,775
Ensembl chr 6:89,483,727...89,504,965
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Psma4
proteasome 20S subunit alpha 4
part_of
ISO
(PMID:17323924)
RGD
PMID:17323924
NCBI chr 8:55,340,431...55,347,893
Ensembl chr 8:55,340,386...55,347,890
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Psma5
proteasome 20S subunit alpha 5
part_of
ISO
(PMID:17323924)
RGD
PMID:17323924
NCBI chr 2:195,896,369...195,919,733
Ensembl chr 2:195,896,365...195,919,731
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Psma6
proteasome 20S subunit alpha 6
part_of
ISO
(PMID:17323924)
RGD
PMID:17323924
NCBI chr 6:72,765,534...72,796,554
Ensembl chr 6:72,765,473...72,796,554
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Psma7
proteasome 20S subunit alpha 7
part_of
ISO
(PMID:17323924)
RGD
PMID:17323924
NCBI chr 3:167,137,279...167,143,626
Ensembl chr 3:167,137,263...167,143,672
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Psmb1
proteasome 20S subunit beta 1
part_of
ISO
(PMID:17323924)
RGD
PMID:17323924
NCBI chr 1:56,442,432...56,463,544
Ensembl chr 1:56,420,618...56,463,560
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Psmb2
proteasome 20S subunit beta 2
part_of
ISO
(PMID:17323924)
RGD
PMID:17323924
NCBI chr 5:138,970,564...139,002,715
Ensembl chr 5:138,970,533...139,021,137
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Psmb3
proteasome 20S subunit beta 3
part_of
ISO
(PMID:17323924)
RGD
PMID:17323924
NCBI chr10:82,697,425...82,704,521
Ensembl chr10:82,697,425...82,704,521
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Psmb4
proteasome 20S subunit beta 4
part_of
ISO
(PMID:17323924), (PMID:26524591)
RGD
PMID:17323924 PMID:26524591
NCBI chr 2:182,442,757...182,445,532
Ensembl chr 2:182,442,756...182,445,746
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Psmb5
proteasome 20S subunit beta 5
part_of
ISO
(PMID:17323924)
RGD
PMID:17323924
NCBI chr15:28,072,772...28,077,367
Ensembl chr15:28,072,781...28,077,440
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Psmb6
proteasome 20S subunit beta 6
part_of
ISO
(PMID:17323924)
RGD
PMID:17323924
NCBI chr10:55,239,256...55,241,586
Ensembl chr10:55,239,241...55,241,586
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Psmb7
proteasome 20S subunit beta 7
part_of
ISO
(PMID:17323924)
RGD
PMID:17323924
NCBI chr 3:22,357,777...22,417,748
Ensembl chr 3:22,354,698...22,417,937
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Psmb8
proteasome 20S subunit beta 8
part_of
ISO
(PMID:26524591)
RGD
PMID:26524591
NCBI chr20:4,652,159...4,655,122
Ensembl chr20:4,652,159...4,655,283
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Psmb9
proteasome 20S subunit beta 9
IDA
RGD
PMID:11295489
RGD:9854635
NCBI chr20:4,667,044...4,672,512
Ensembl chr20:4,666,046...4,672,512
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Psmc1
proteasome 26S subunit, ATPase 1
part_of
ISO IEA
(PMID:17323924) GO_REF:0000043
RGD UniProt
PMID:17323924
GO_REF:0000043
NCBI chr 6:119,392,833...119,405,233
Ensembl chr 6:119,392,855...119,410,123
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Psmc2
proteasome 26S subunit, ATPase 2
part_of
ISO IEA
(PMID:17323924), (PMID:9295362), (PMID:9464850) GO_REF:0000107
RGD Ensembl
PMID:9295362 PMID:9464850 PMID:17323924
GO_REF:0000107
NCBI chr 4:13,255,850...13,270,160
Ensembl chr 4:13,255,856...13,270,185
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Psmc3
proteasome 26S subunit, ATPase 3
part_of
ISO IEA
(PMID:17323924), (PMID:9464850) GO_REF:0000107 GO_REF:0000043
RGD Ensembl UniProt
PMID:9464850 PMID:17323924
GO_REF:0000043 GO_REF:0000107
NCBI chr 3:77,031,825...77,037,207
Ensembl chr 3:77,031,825...77,043,358
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Psmc4
proteasome 26S subunit, ATPase 4
part_of
ISO
(PMID:17323924)
RGD
PMID:17323924
NCBI chr 1:83,349,127...83,357,497
Ensembl chr 1:83,348,592...83,357,494
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Psmc5
proteasome 26S subunit, ATPase 5
part_of
ISO ISS
(PMID:17323924), (PMID:9464850) GO_REF:0000024
RGD UniProt
PMID:9464850 PMID:17323924
GO_REF:0000024
NCBI chr10:91,187,763...91,193,697
Ensembl chr10:91,187,743...91,213,135
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Psmc6
proteasome 26S subunit, ATPase 6
part_of
ISO
(PMID:17323924), (PMID:9464850)
RGD
PMID:9464850 PMID:17323924
NCBI chr15:18,542,585...18,564,057
Ensembl chr15:18,542,563...18,564,055
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Psmd1
proteasome 26S subunit, non-ATPase 1
part_of
ISO
(PMID:17323924) (PMID:16973439)
RGD
PMID:16973439 PMID:17323924
NCBI chr 9:86,709,714...86,785,213
Ensembl chr 9:86,709,947...86,785,211
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Psmd10
proteasome 26S subunit, non-ATPase 10
part_of
ISO
(PMID:17323924)
RGD
PMID:17323924
NCBI chr X:104,656,809...104,665,122
Ensembl chr X:104,656,812...104,665,097
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Psmd11
proteasome 26S subunit, non-ATPase 11
part_of
ISO IEA
(PMID:17323924) GO_REF:0000043
RGD UniProt
PMID:17323924
GO_REF:0000043
NCBI chr10:65,433,853...65,479,774
Ensembl chr10:65,433,853...65,479,766
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Psmd12
proteasome 26S subunit, non-ATPase 12
part_of
ISO
(PMID:17323924)
RGD
PMID:17323924
NCBI chr10:92,438,223...92,457,323
Ensembl chr10:92,438,298...92,457,287
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Psmd13
proteasome 26S subunit, non-ATPase 13
part_of
ISO
(PMID:17323924)
RGD
PMID:17323924
NCBI chr 1:195,964,617...195,976,895
Ensembl chr 1:195,964,138...195,976,905
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Psmd14
proteasome 26S subunit, non-ATPase 14
part_of
ISO
(PMID:17323924) (PMID:16973439)
RGD
PMID:16973439 PMID:17323924
NCBI chr 3:46,254,338...46,347,076
Ensembl chr 3:46,254,330...46,347,076
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Psmd2
proteasome 26S subunit ubiquitin receptor, non-ATPase 2
part_of
ISO
(PMID:17323924) (PMID:16973439)
RGD
PMID:16973439 PMID:17323924
NCBI chr11:80,248,364...80,258,991
Ensembl chr11:80,248,364...80,259,043
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Psmd3
proteasome 26S subunit, non-ATPase 3
part_of
ISO
(PMID:17323924)
RGD
PMID:17323924
NCBI chr10:83,643,473...83,655,620
Ensembl chr10:83,643,647...83,655,618
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Psmd4
proteasome 26S subunit ubiquitin receptor, non-ATPase 4
part_of
ISO
(PMID:17323924)
RGD
PMID:17323924
NCBI chr 2:182,598,934...182,608,250
Ensembl chr 2:182,598,934...182,608,194
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Psmd6
proteasome 26S subunit, non-ATPase 6
part_of
ISO
(PMID:17323924)
RGD
PMID:17323924
NCBI chr15:11,102,174...11,112,180
Ensembl chr15:11,102,180...11,205,620
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Psmd7
proteasome 26S subunit, non-ATPase 7
part_of
ISO IBA
(PMID:17323924) GO_REF:0000033
RGD GO_Central
PMID:17323924
GO_REF:0000033
NCBI chr19:35,494,317...35,501,605
Ensembl chr19:35,494,316...35,501,588
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Psmd8
proteasome 26S subunit, non-ATPase 8
part_of
ISO
(PMID:17323924)
RGD
PMID:17323924
NCBI chr 1:84,474,377...84,481,209
Ensembl chr 1:84,474,378...84,481,209
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Psmf1
proteasome inhibitor subunit 1
part_of
IEA
GO_REF:0000043
UniProt
GO_REF:0000043
NCBI chr 3:140,235,371...140,260,546
Ensembl chr 3:140,235,373...140,260,546
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Rad23a
RAD23 homolog A, nucleotide excision repair protein
part_of
IEA
GO_REF:0000043
UniProt
GO_REF:0000043
NCBI chr19:23,313,563...23,320,702
Ensembl chr19:23,314,797...23,320,695
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Rad23b
RAD23 homolog B, nucleotide excision repair protein
part_of
IEA
GO_REF:0000043
UniProt
GO_REF:0000043
NCBI chr 5:70,090,232...70,128,340
Ensembl chr 5:70,090,232...70,128,319
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Sem1
SEM1 26S proteasome subunit
part_of
ISO IBA
(PMID:15117943), (PMID:17323924) GO_REF:0000033
RGD GO_Central
PMID:15117943 PMID:17323924
GO_REF:0000033
NCBI chr 4:34,671,848...34,692,005
Ensembl chr 4:34,671,848...34,692,005
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Slc10a2
solute carrier family 10 member 2
IDA
RGD
PMID:15304498
RGD:1624187
NCBI chr16:84,386,528...84,409,475
Ensembl chr16:84,374,862...84,409,475
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Txnl1
thioredoxin-like 1
part_of
IEA
GO_REF:0000043
UniProt
GO_REF:0000043
NCBI chr18:57,125,362...57,155,167
Ensembl chr18:57,125,369...57,165,266
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Ube3a
ubiquitin protein ligase E3A
part_of
IEA
GO_REF:0000043
UniProt
GO_REF:0000043
NCBI chr 1:110,070,260...110,161,675
Ensembl chr 1:110,070,480...110,157,250
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Ubqln1
ubiquilin 1
part_of
IEA
GO_REF:0000043
UniProt
GO_REF:0000043
NCBI chr17:6,439,002...6,477,096
Ensembl chr17:6,439,002...6,477,090
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Ubr1
ubiquitin protein ligase E3 component n-recognin 1
part_of
ISO
SGD:S000003416 (PMID:11689692)
RGD
PMID:11689692
NCBI chr 3:107,813,721...107,921,701
Ensembl chr 3:107,811,392...107,922,204
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Usp14
ubiquitin specific peptidase 14
part_of
IEA
GO_REF:0000043
UniProt
GO_REF:0000043
NCBI chr18:1,018,111...1,057,727
Ensembl chr18:1,019,400...1,057,519
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Vcp
valosin-containing protein
part_of
ISO
(PMID:9452483)
RGD
PMID:9452483
NCBI chr 5:57,210,167...57,229,571
Ensembl chr 5:57,210,168...57,229,571
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Zfand2a
zinc finger AN1-type containing 2A
part_of
ISO
(PMID:16973439)
RGD
PMID:16973439
NCBI chr12:15,161,601...15,172,679
Ensembl chr12:15,161,639...15,172,677
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Zfand2b
zinc finger AN1-type containing 2B
part_of located_in
ISO
(MGI:6092631|PMID:18467495) (PMID:18467495) (PMID:26876100)
RGD
PMID:18467495 PMID:26876100
MGI:6092631
NCBI chr 9:76,665,466...76,668,447
Ensembl chr 9:76,665,546...76,668,445
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Ide
insulin degrading enzyme
IDA
RGD
PMID:9000694
RGD:1627643
NCBI chr 1:235,002,984...235,102,448
Ensembl chr 1:234,995,351...235,102,440
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Psmc4
proteasome 26S subunit, ATPase 4
IDA
RGD
PMID:18986984
RGD:7242199
NCBI chr 1:83,349,127...83,357,497
Ensembl chr 1:83,348,592...83,357,494
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Psmc5
proteasome 26S subunit, ATPase 5
IDA
RGD
PMID:18986984
RGD:7242199
NCBI chr10:91,187,763...91,193,697
Ensembl chr10:91,187,743...91,213,135
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Psmc6
proteasome 26S subunit, ATPase 6
part_of
IDA IBA
GO_REF:0000033
GO_Central RGD
PMID:18986984
GO_REF:0000033, RGD:7242199
NCBI chr15:18,542,585...18,564,057
Ensembl chr15:18,542,563...18,564,055
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Psmd14
proteasome 26S subunit, non-ATPase 14
IDA
RGD
PMID:19609968
RGD:9480236
NCBI chr 3:46,254,338...46,347,076
Ensembl chr 3:46,254,330...46,347,076
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Ubqln4
ubiquilin 4
located_in
ISO
(PMID:15280365)
RGD
PMID:15280365
NCBI chr 2:174,012,726...174,028,062
Ensembl chr 2:174,012,777...174,028,059
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Uchl5
ubiquitin C-terminal hydrolase L5
IDA
RGD
PMID:19609968
RGD:9480236
NCBI chr13:55,520,722...55,556,503
Ensembl chr13:55,520,729...55,556,502
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Psmc5
proteasome 26S subunit, ATPase 5
IDA
RGD
PMID:8710853
RGD:6771234
NCBI chr10:91,187,763...91,193,697
Ensembl chr10:91,187,743...91,213,135
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Ubqln4
ubiquilin 4
located_in
ISO
(PMID:15280365)
RGD
PMID:15280365
NCBI chr 2:174,012,726...174,028,062
Ensembl chr 2:174,012,777...174,028,059
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Psmc1
proteasome 26S subunit, ATPase 1
part_of
ISS ISO
GO_REF:0000024 (PMID:16857966) (MGI:5442509|PMID:16857966)
UniProt RGD
PMID:16857966
GO_REF:0000024 MGI:5442509
NCBI chr 6:119,392,833...119,405,233
Ensembl chr 6:119,392,855...119,410,123
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Psmc2
proteasome 26S subunit, ATPase 2
part_of
ISS ISO
GO_REF:0000024 (PMID:16857966) (MGI:5442509|PMID:16857966)
UniProt RGD
PMID:16857966
GO_REF:0000024 MGI:5442509
NCBI chr 4:13,255,850...13,270,160
Ensembl chr 4:13,255,856...13,270,185
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Psmc3
proteasome 26S subunit, ATPase 3
part_of
ISS ISO
GO_REF:0000024 (MGI:5442509|PMID:16857966)
UniProt RGD
PMID:16857966
GO_REF:0000024 MGI:5442509
NCBI chr 3:77,031,825...77,037,207
Ensembl chr 3:77,031,825...77,043,358
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Psmc4
proteasome 26S subunit, ATPase 4
part_of
ISS ISO
GO_REF:0000024 (PMID:16857966) (MGI:5442509|PMID:16857966)
UniProt RGD
PMID:16857966
GO_REF:0000024 MGI:5442509
NCBI chr 1:83,349,127...83,357,497
Ensembl chr 1:83,348,592...83,357,494
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Psmc5
proteasome 26S subunit, ATPase 5
part_of
ISS ISO
GO_REF:0000024 (PMID:16857966) (MGI:5442509|PMID:16857966)
UniProt RGD
PMID:16857966
GO_REF:0000024 MGI:5442509
NCBI chr10:91,187,763...91,193,697
Ensembl chr10:91,187,743...91,213,135
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Psmc6
proteasome 26S subunit, ATPase 6
part_of
ISO
(MGI:5442509|PMID:16857966) (PMID:16857966)
RGD
PMID:16857966
MGI:5442509
NCBI chr15:18,542,585...18,564,057
Ensembl chr15:18,542,563...18,564,055
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Psmd1
proteasome 26S subunit, non-ATPase 1
part_of
ISS ISO
GO_REF:0000024 (PMID:16857966) (MGI:5442509|PMID:16857966)
UniProt RGD
PMID:16857966
GO_REF:0000024 MGI:5442509
NCBI chr 9:86,709,714...86,785,213
Ensembl chr 9:86,709,947...86,785,211
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Psmd11
proteasome 26S subunit, non-ATPase 11
part_of
ISO ISS
(PMID:22972301) (PMID:16857966) (MGI:5442509|PMID:16857966) GO_REF:0000024
RGD UniProt
PMID:16857966 PMID:22972301
GO_REF:0000024 MGI:5442509
NCBI chr10:65,433,853...65,479,774
Ensembl chr10:65,433,853...65,479,766
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Psmd12
proteasome 26S subunit, non-ATPase 12
part_of
ISO
(MGI:5442509|PMID:16857966) (PMID:16857966)
RGD
PMID:16857966
MGI:5442509
NCBI chr10:92,438,223...92,457,323
Ensembl chr10:92,438,298...92,457,287
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Psmd13
proteasome 26S subunit, non-ATPase 13
part_of
ISS ISO
GO_REF:0000024 (PMID:16857966) (MGI:5442509|PMID:16857966)
UniProt RGD
PMID:16857966
GO_REF:0000024 MGI:5442509
NCBI chr 1:195,964,617...195,976,895
Ensembl chr 1:195,964,138...195,976,905
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Psmd14
proteasome 26S subunit, non-ATPase 14
part_of
ISO
(MGI:5442509|PMID:16857966) (PMID:16857966)
RGD
PMID:16857966
MGI:5442509
NCBI chr 3:46,254,338...46,347,076
Ensembl chr 3:46,254,330...46,347,076
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Psmd2
proteasome 26S subunit ubiquitin receptor, non-ATPase 2
part_of
ISS ISO
GO_REF:0000024 (PMID:16857966) (MGI:5442509|PMID:16857966)
UniProt RGD
PMID:16857966
GO_REF:0000024 MGI:5442509
NCBI chr11:80,248,364...80,258,991
Ensembl chr11:80,248,364...80,259,043
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Psmd3
proteasome 26S subunit, non-ATPase 3
part_of
ISO
(MGI:5442509|PMID:16857966) (PMID:16857966)
RGD
PMID:16857966
MGI:5442509
NCBI chr10:83,643,473...83,655,620
Ensembl chr10:83,643,647...83,655,618
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Psmd4
proteasome 26S subunit ubiquitin receptor, non-ATPase 4
part_of
ISO
(MGI:5442509|PMID:16857966) (PMID:16857966)
RGD
PMID:16857966
MGI:5442509
NCBI chr 2:182,598,934...182,608,250
Ensembl chr 2:182,598,934...182,608,194
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Psmd5
proteasome 26S subunit, non-ATPase 5
part_of
ISO
(MGI:5442509|PMID:16857966)
RGD
PMID:16857966
MGI:5442509
NCBI chr 3:18,133,715...18,151,000
Ensembl chr 3:18,133,717...18,151,029
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Psmd6
proteasome 26S subunit, non-ATPase 6
part_of
ISO
(MGI:5442509|PMID:16857966)
RGD
PMID:16857966
MGI:5442509
NCBI chr15:11,102,174...11,112,180
Ensembl chr15:11,102,180...11,205,620
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Psmd8
proteasome 26S subunit, non-ATPase 8
part_of
ISO
(MGI:5442509|PMID:16857966) (PMID:16857966)
RGD
PMID:16857966
MGI:5442509
NCBI chr 1:84,474,377...84,481,209
Ensembl chr 1:84,474,378...84,481,209
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AABR07062753.1
part_of
IEA
GO_REF:0000002
InterPro
GO_REF:0000002
Ensembl chr 6:11,713,966...11,714,477
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G6pc1
glucose-6-phosphatase catalytic subunit 1
part_of
IEA
GO_REF:0000002
InterPro
GO_REF:0000002
NCBI chr10:86,307,400...86,318,766
Ensembl chr10:86,257,668...86,333,804
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LOC100362709
proteasome activator complex subunit 2-like
part_of
IEA
GO_REF:0000002
InterPro
GO_REF:0000002
NCBI chr 1:34,722,205...34,723,042
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Psme1
proteasome activator subunit 1
part_of
IEA
GO_REF:0000002
InterPro
GO_REF:0000002
NCBI chr15:29,069,012...29,071,888
Ensembl chr15:29,068,729...29,071,890
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Psme1-ps1
proteasome activator subunit 1, pseudogene 1
part_of
IEA
GO_REF:0000002
InterPro
GO_REF:0000002
NCBI chr17:50,174,233...50,175,012
Ensembl chr17:50,174,186...50,175,160
G
Psme2
proteasome activator subunit 2
part_of
IEA
GO_REF:0000002
InterPro
GO_REF:0000002
NCBI chr15:29,078,495...29,082,794
Ensembl chr15:29,078,500...29,082,946
G
Psme3
proteasome activator subunit 3
part_of
IEA
GO_REF:0000002
InterPro
GO_REF:0000002
NCBI chr10:86,257,672...86,265,458
Ensembl chr10:86,257,668...86,333,804
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Ctrl
chymotrypsin-like
part_of
IEA
GO_REF:0000002
InterPro
GO_REF:0000002
NCBI chr19:33,827,279...33,829,129
Ensembl chr19:33,827,229...33,833,626
G
ENSRNOG00000062555
part_of
IEA
GO_REF:0000002
InterPro
GO_REF:0000002
Ensembl chr 1:224,032,844...224,034,987
G
ENSRNOG00000069501
part_of
IEA
GO_REF:0000002
InterPro
GO_REF:0000002
Ensembl chr 7:96,706,207...96,707,107
G
ENSRNOG00000070269
part_of
IEA
GO_REF:0000002
InterPro
GO_REF:0000002
Ensembl chr X:51,562,982...51,565,263
G
ENSRNOG00000070633
part_of
IEA
GO_REF:0000002
InterPro
GO_REF:0000002
Ensembl chr15:24,257,489...24,258,627
G
Psma1
proteasome 20S subunit alpha 1
part_of
ISS ISO
GO_REF:0000024 (PMID:16857966), (PMID:20498273) (MGI:5442509|PMID:16857966), (PMID:20498273) (PMID:34711951)
UniProt RGD
PMID:16857966 PMID:20498273 PMID:34711951
GO_REF:0000024 MGI:5442509
NCBI chr 1:168,442,421...168,453,440
Ensembl chr 1:168,440,936...168,453,472
G
Psma2
proteasome 20S subunit alpha 2
part_of
ISO ISS
(PMID:34711951) (PMID:16857966), (PMID:20498273) (MGI:5442509|PMID:16857966), (PMID:20498273) GO_REF:0000024
RGD UniProt
PMID:16857966 PMID:20498273 PMID:34711951
GO_REF:0000024 MGI:5442509
NCBI chr17:50,554,536...50,564,923
Ensembl chr17:50,551,924...50,564,938
G
Psma3
proteasome 20S subunit alpha 3
part_of
ISO ISS
(PMID:19208651), (PMID:34711951) (PMID:16857966), (PMID:20498273) (MGI:5442509|PMID:16857966), (PMID:20498273) GO_REF:0000024
RGD UniProt
PMID:16857966 PMID:19208651 PMID:20498273 PMID:34711951
GO_REF:0000024 MGI:5442509
NCBI chr 6:89,483,741...89,503,775
Ensembl chr 6:89,483,727...89,504,965
G
Psma4
proteasome 20S subunit alpha 4
part_of
ISS ISO
GO_REF:0000024 (PMID:16857966), (PMID:20498273) (MGI:5442509|PMID:16857966), (PMID:20498273) (PMID:34711951)
UniProt RGD
PMID:16857966 PMID:20498273 PMID:34711951
GO_REF:0000024 MGI:5442509
NCBI chr 8:55,340,431...55,347,893
Ensembl chr 8:55,340,386...55,347,890
G
Psma5
proteasome 20S subunit alpha 5
part_of
ISS ISO
GO_REF:0000024 (MGI:5442509|PMID:16857966), (PMID:17540904), (PMID:20498273) (PMID:34711951)
UniProt RGD
PMID:16857966 PMID:17540904 PMID:20498273 PMID:34711951
GO_REF:0000024 MGI:5442509
NCBI chr 2:195,896,369...195,919,733
Ensembl chr 2:195,896,365...195,919,731
G
Psma6
proteasome 20S subunit alpha 6
part_of
ISS ISO
GO_REF:0000024 (MGI:5442509|PMID:16857966), (PMID:20498273) (PMID:34711951)
UniProt RGD
PMID:16857966 PMID:20498273 PMID:34711951
GO_REF:0000024 MGI:5442509
NCBI chr 6:72,765,534...72,796,554
Ensembl chr 6:72,765,473...72,796,554
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Psma6-ps5
proteasome 20S subunit alpha 6, pseudogene 5
part_of
IEA
GO_REF:0000002
InterPro
GO_REF:0000002
NCBI chr20:52,127,531...52,129,095
G
Psma7
proteasome 20S subunit alpha 7
part_of
ISS ISO
GO_REF:0000024 (PMID:16857966), (PMID:20498273) (MGI:5442509|PMID:16857966), (PMID:20498273) (PMID:34711951)
UniProt RGD
PMID:16857966 PMID:20498273 PMID:34711951
GO_REF:0000024 MGI:5442509
NCBI chr 3:167,137,279...167,143,626
Ensembl chr 3:167,137,263...167,143,672
G
Psmb1
proteasome 20S subunit beta 1
part_of
IEA ISO IBA ISS
GO_REF:0000107 (PMID:16857966), (PMID:17540904), (PMID:20498273) (MGI:5442509|PMID:16857966), (PMID:17540904), (PMID:20498273) GO_REF:0000033 GO_REF:0000002 (PMID:34711951) GO_REF:0000024
Ensembl RGD GO_Central InterPro UniProt
PMID:16857966 PMID:17540904 PMID:20498273 PMID:34711951
GO_REF:0000002 GO_REF:0000024 GO_REF:0000033 GO_REF:0000107 MGI:5442509
NCBI chr 1:56,442,432...56,463,544
Ensembl chr 1:56,420,618...56,463,560
G
Psmb10
proteasome 20S subunit beta 10
part_of
ISS ISO IEA
GO_REF:0000024 (PMID:16857966), (PMID:17540904) (MGI:5442509|PMID:16857966), (PMID:17540904) GO_REF:0000002
UniProt RGD InterPro
PMID:16857966 PMID:17540904
GO_REF:0000002 GO_REF:0000024 MGI:5442509
NCBI chr19:33,830,958...33,833,442
Ensembl chr19:33,827,229...33,833,626
G
Psmb11
proteasome subunit beta 11
part_of
ISO
(PMID:17540904)
RGD
PMID:17540904
NCBI chr15:28,084,736...28,085,655
G
Psmb2
proteasome 20S subunit beta 2
part_of
ISO ISS
(PMID:34711951) (PMID:16857966), (PMID:17540904), (PMID:20498273) (MGI:5442509|PMID:16857966), (PMID:17540904), (PMID:20498273) GO_REF:0000024
RGD UniProt
PMID:16857966 PMID:17540904 PMID:20498273 PMID:34711951
GO_REF:0000024 MGI:5442509
NCBI chr 5:138,970,564...139,002,715
Ensembl chr 5:138,970,533...139,021,137
G
Psmb3
proteasome 20S subunit beta 3
part_of
ISS ISO
GO_REF:0000024 (PMID:10436176), (PMID:16857966), (PMID:20498273) (MGI:5442509|PMID:16857966), (PMID:10436176), (PMID:20498273) (PMID:34711951)
UniProt RGD
PMID:10436176 PMID:16857966 PMID:20498273 PMID:34711951
GO_REF:0000024 MGI:5442509
NCBI chr10:82,697,425...82,704,521
Ensembl chr10:82,697,425...82,704,521
G
Psmb4
proteasome 20S subunit beta 4
part_of
ISS ISO IEA
GO_REF:0000024 (MGI:5442509|PMID:16857966), (PMID:20498273) GO_REF:0000107 (PMID:34711951)
UniProt RGD Ensembl
PMID:16857966 PMID:20498273 PMID:34711951
GO_REF:0000024 GO_REF:0000107 MGI:5442509
NCBI chr 2:182,442,757...182,445,532
Ensembl chr 2:182,442,756...182,445,746
G
Psmb5
proteasome 20S subunit beta 5
part_of
IEA ISO ISS
GO_REF:0000107 (PMID:15198663), (PMID:16857966), (PMID:17540904), (PMID:20498273) (MGI:5442509|PMID:16857966), (PMID:15198663), (PMID:17540904), (PMID:20498273) (PMID:34711951) GO_REF:0000024
Ensembl RGD UniProt
PMID:15198663 PMID:16857966 PMID:17540904 PMID:20498273 PMID:34711951
GO_REF:0000024 GO_REF:0000107 MGI:5442509
NCBI chr15:28,072,772...28,077,367
Ensembl chr15:28,072,781...28,077,440
G
Psmb6
proteasome 20S subunit beta 6
part_of
ISS ISO
GO_REF:0000024 (PMID:16857966), (PMID:20498273) (MGI:5442509|PMID:16857966), (PMID:20498273) (PMID:34711951)
UniProt RGD
PMID:16857966 PMID:20498273 PMID:34711951
GO_REF:0000024 MGI:5442509
NCBI chr10:55,239,256...55,241,586
Ensembl chr10:55,239,241...55,241,586
G
Psmb6l1
proteasome 20S subunit beta 6 like 1
part_of
ISS
GO_REF:0000024
UniProt
GO_REF:0000024
NCBI chr 2:93,833,914...93,834,630
Ensembl chr 2:93,833,830...93,834,635
G
Psmb7
proteasome 20S subunit beta 7
part_of
ISS ISO IEA
GO_REF:0000024 (PMID:16857966), (PMID:20498273) (MGI:5442509|PMID:16857966), (PMID:20498273) GO_REF:0000107 (PMID:34711951)
UniProt RGD Ensembl
PMID:16857966 PMID:20498273 PMID:34711951
GO_REF:0000024 GO_REF:0000107 MGI:5442509
NCBI chr 3:22,357,777...22,417,748
Ensembl chr 3:22,354,698...22,417,937
G
Psmb8
proteasome 20S subunit beta 8
part_of
TAS ISO IEA ISS
(PMID:16857966), (PMID:17540904) (MGI:5442509|PMID:16857966), (PMID:17540904) GO_REF:0000107 GO_REF:0000024
RGD Ensembl UniProt
PMID:16857966 PMID:17540904 PMID:1451788
GO_REF:0000024 GO_REF:0000107 MGI:5442509 , RGD:70234
NCBI chr20:4,652,159...4,655,122
Ensembl chr20:4,652,159...4,655,283
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Psmb9
proteasome 20S subunit beta 9
part_of
IEA ISO
GO_REF:0000002 (MGI:5442509|PMID:16857966), (PMID:17540904)
InterPro RGD
PMID:16857966 PMID:17540904
GO_REF:0000002 MGI:5442509
NCBI chr20:4,667,044...4,672,512
Ensembl chr20:4,666,046...4,672,512
G
ENSRNOG00000066197
part_of
IEA
GO_REF:0000002
InterPro
GO_REF:0000002
Ensembl chr 8:84,687,138...84,687,667
G
ENSRNOG00000066438
part_of
IEA
GO_REF:0000104
UniProt
GO_REF:0000104
Ensembl chr 4:27,837,452...27,838,144
G
ENSRNOG00000069956
part_of
IEA
GO_REF:0000002
InterPro
GO_REF:0000002
Ensembl chr12:3,770,126...3,770,382
G
Psma1
proteasome 20S subunit alpha 1
part_of
ISS IBA
GO_REF:0000024 GO_REF:0000033
UniProt GO_Central
GO_REF:0000024 GO_REF:0000033
NCBI chr 1:168,442,421...168,453,440
Ensembl chr 1:168,440,936...168,453,472
G
Psma2
proteasome 20S subunit alpha 2
part_of
ISS IBA IEA
GO_REF:0000024 GO_REF:0000033 GO_REF:0000104
UniProt GO_Central
GO_REF:0000024 GO_REF:0000033 GO_REF:0000104
NCBI chr17:50,554,536...50,564,923
Ensembl chr17:50,551,924...50,564,938
G
Psma3
proteasome 20S subunit alpha 3
part_of
ISS IBA IEA
GO_REF:0000024 GO_REF:0000033 GO_REF:0000104
UniProt GO_Central
GO_REF:0000024 GO_REF:0000033 GO_REF:0000104
NCBI chr 6:89,483,741...89,503,775
Ensembl chr 6:89,483,727...89,504,965
G
Psma4
proteasome 20S subunit alpha 4
part_of
ISO IBA ISS
(PMID:30133132) GO_REF:0000033 GO_REF:0000024
RGD GO_Central UniProt
PMID:30133132
GO_REF:0000024 GO_REF:0000033
NCBI chr 8:55,340,431...55,347,893
Ensembl chr 8:55,340,386...55,347,890
G
Psma5
proteasome 20S subunit alpha 5
part_of
ISO IBA ISS IEA
(PMID:30133132) GO_REF:0000033 GO_REF:0000024 GO_REF:0000104
RGD GO_Central UniProt
PMID:30133132
GO_REF:0000024 GO_REF:0000033 GO_REF:0000104
NCBI chr 2:195,896,369...195,919,733
Ensembl chr 2:195,896,365...195,919,731
G
Psma6
proteasome 20S subunit alpha 6
part_of
ISS IBA IEA ISO
GO_REF:0000024 GO_REF:0000033 GO_REF:0000104 (PMID:30133132), (PMID:9096325)
BHF-UCL GO_Central UniProt RGD
PMID:9096325 PMID:30133132
GO_REF:0000024 GO_REF:0000033 GO_REF:0000104
NCBI chr 6:72,765,534...72,796,554
Ensembl chr 6:72,765,473...72,796,554
G
Psma7
proteasome 20S subunit alpha 7
part_of
ISO IBA IEA ISS
(PMID:30133132) GO_REF:0000033 GO_REF:0000104 GO_REF:0000024
RGD GO_Central UniProt
PMID:30133132
GO_REF:0000024 GO_REF:0000033 GO_REF:0000104
NCBI chr 3:167,137,279...167,143,626
Ensembl chr 3:167,137,263...167,143,672
G
Psma8
proteasome 20S subunit alpha 8
part_of
IBA
GO_REF:0000033
GO_Central
GO_REF:0000033
NCBI chr18:5,863,594...5,928,226
Ensembl chr18:5,863,608...5,928,226
G
Psmb1
proteasome 20S subunit beta 1
part_of
ISS
GO_REF:0000024
UniProt
GO_REF:0000024
NCBI chr 1:56,442,432...56,463,544
Ensembl chr 1:56,420,618...56,463,560
G
Psmb10
proteasome 20S subunit beta 10
part_of
IBA
GO_REF:0000033
GO_Central
GO_REF:0000033
NCBI chr19:33,830,958...33,833,442
Ensembl chr19:33,827,229...33,833,626
G
Psmb2
proteasome 20S subunit beta 2
part_of
ISS IBA
GO_REF:0000024 GO_REF:0000033
UniProt GO_Central
GO_REF:0000024 GO_REF:0000033
NCBI chr 5:138,970,564...139,002,715
Ensembl chr 5:138,970,533...139,021,137
G
Psmb3
proteasome 20S subunit beta 3
part_of
ISS IBA IEA
GO_REF:0000024 GO_REF:0000033 GO_REF:0000002
UniProt GO_Central InterPro
GO_REF:0000002 GO_REF:0000024 GO_REF:0000033
NCBI chr10:82,697,425...82,704,521
Ensembl chr10:82,697,425...82,704,521
G
Psmb3l2
proteasome 20S subunit beta 3 like 2
part_of
IBA
GO_REF:0000033
GO_Central
GO_REF:0000033
NCBI chr 3:118,672,316...118,672,933
Ensembl chr 3:118,672,233...118,672,933
G
Psmb4
proteasome 20S subunit beta 4
part_of
ISS IBA
GO_REF:0000024 GO_REF:0000033
UniProt GO_Central
GO_REF:0000024 GO_REF:0000033
NCBI chr 2:182,442,757...182,445,532
Ensembl chr 2:182,442,756...182,445,746
G
Psmb5
proteasome 20S subunit beta 5
part_of
ISS IBA
GO_REF:0000024 GO_REF:0000033
UniProt GO_Central
GO_REF:0000024 GO_REF:0000033
NCBI chr15:28,072,772...28,077,367
Ensembl chr15:28,072,781...28,077,440
G
Psmb6
proteasome 20S subunit beta 6
part_of
ISS IBA
GO_REF:0000024 GO_REF:0000033
UniProt GO_Central
GO_REF:0000024 GO_REF:0000033
NCBI chr10:55,239,256...55,241,586
Ensembl chr10:55,239,241...55,241,586
G
Psmb6l1
proteasome 20S subunit beta 6 like 1
part_of
ISS IBA
GO_REF:0000024 GO_REF:0000033
UniProt GO_Central
GO_REF:0000024 GO_REF:0000033
NCBI chr 2:93,833,914...93,834,630
Ensembl chr 2:93,833,830...93,834,635
G
Psmb7
proteasome 20S subunit beta 7
part_of
ISS IBA
GO_REF:0000024 GO_REF:0000033
UniProt GO_Central
GO_REF:0000024 GO_REF:0000033
NCBI chr 3:22,357,777...22,417,748
Ensembl chr 3:22,354,698...22,417,937
G
Psmb8
proteasome 20S subunit beta 8
part_of
ISS IBA
GO_REF:0000024 GO_REF:0000033
UniProt GO_Central
GO_REF:0000024 GO_REF:0000033
NCBI chr20:4,652,159...4,655,122
Ensembl chr20:4,652,159...4,655,283
G
Psmb9
proteasome 20S subunit beta 9
part_of
ISS IBA
GO_REF:0000024 GO_REF:0000033
UniProt GO_Central
GO_REF:0000024 GO_REF:0000033
NCBI chr20:4,667,044...4,672,512
Ensembl chr20:4,666,046...4,672,512
G
ENSRNOG00000065937
part_of
IEA
GO_REF:0000002
InterPro
GO_REF:0000002
Ensembl chr 3:45,456,139...45,457,335
G
Psmc1
proteasome 26S subunit, ATPase 1
part_of
ISO IEA
(PMID:20498273) GO_REF:0000107
RGD Ensembl
PMID:20498273
GO_REF:0000107
NCBI chr 6:119,392,833...119,405,233
Ensembl chr 6:119,392,855...119,410,123
G
Psmc5
proteasome 26S subunit, ATPase 5
part_of
ISO IEA
(PMID:20498273) GO_REF:0000107
RGD Ensembl
PMID:20498273
GO_REF:0000107
NCBI chr10:91,187,763...91,193,697
Ensembl chr10:91,187,743...91,213,135
G
Psmd11
proteasome 26S subunit, non-ATPase 11
part_of
ISO IEA
(PMID:20498273) GO_REF:0000107
RGD Ensembl
PMID:20498273
GO_REF:0000107
NCBI chr10:65,433,853...65,479,774
Ensembl chr10:65,433,853...65,479,766
G
Psmd12
proteasome 26S subunit, non-ATPase 12
part_of
ISO
(PMID:20498273)
RGD
PMID:20498273 PMID:9426256
RGD:8693626
NCBI chr10:92,438,223...92,457,323
Ensembl chr10:92,438,298...92,457,287
G
Psmd13
proteasome 26S subunit, non-ATPase 13
part_of
ISO IEA
(PMID:10225435) GO_REF:0000107
RGD Ensembl
PMID:10225435
GO_REF:0000107
NCBI chr 1:195,964,617...195,976,895
Ensembl chr 1:195,964,138...195,976,905
G
Psmd2
proteasome 26S subunit ubiquitin receptor, non-ATPase 2
part_of
ISO
(PMID:20498273)
RGD
PMID:20498273
NCBI chr11:80,248,364...80,258,991
Ensembl chr11:80,248,364...80,259,043
G
Psmd3
proteasome 26S subunit, non-ATPase 3
part_of
ISO
(PMID:20498273)
RGD
PMID:20498273
NCBI chr10:83,643,473...83,655,620
Ensembl chr10:83,643,647...83,655,618
G
Psmd6
proteasome 26S subunit, non-ATPase 6
part_of
IBA
GO_REF:0000033
GO_Central
GO_REF:0000033
NCBI chr15:11,102,174...11,112,180
Ensembl chr15:11,102,180...11,205,620
G
Psmd7
proteasome 26S subunit, non-ATPase 7
part_of
ISO
(PMID:20498273)
RGD
PMID:20498273
NCBI chr19:35,494,317...35,501,605
Ensembl chr19:35,494,316...35,501,588
G
Psmc1
proteasome 26S subunit, ATPase 1
part_of
IBA
GO_REF:0000033
GO_Central
GO_REF:0000033
NCBI chr 6:119,392,833...119,405,233
Ensembl chr 6:119,392,855...119,410,123
G
Psmc2
proteasome 26S subunit, ATPase 2
part_of
IBA
GO_REF:0000033
GO_Central
GO_REF:0000033
NCBI chr 4:13,255,850...13,270,160
Ensembl chr 4:13,255,856...13,270,185
G
Psmc3
proteasome 26S subunit, ATPase 3
part_of
IBA
GO_REF:0000033
GO_Central
GO_REF:0000033
NCBI chr 3:77,031,825...77,037,207
Ensembl chr 3:77,031,825...77,043,358
G
Psmc4
proteasome 26S subunit, ATPase 4
part_of
IBA
GO_REF:0000033
GO_Central
GO_REF:0000033
NCBI chr 1:83,349,127...83,357,497
Ensembl chr 1:83,348,592...83,357,494
G
Psmc5
proteasome 26S subunit, ATPase 5
part_of
IBA
GO_REF:0000033
GO_Central
GO_REF:0000033
NCBI chr10:91,187,763...91,193,697
Ensembl chr10:91,187,743...91,213,135
G
Psmc6
proteasome 26S subunit, ATPase 6
part_of
IBA
GO_REF:0000033
GO_Central
GO_REF:0000033
NCBI chr15:18,542,585...18,564,057
Ensembl chr15:18,542,563...18,564,055
G
Psmd1
proteasome 26S subunit, non-ATPase 1
part_of
IEA IBA
GO_REF:0000104 GO_REF:0000033
UniProt GO_Central
GO_REF:0000033 GO_REF:0000104
NCBI chr 9:86,709,714...86,785,213
Ensembl chr 9:86,709,947...86,785,211
G
Psmd10
proteasome 26S subunit, non-ATPase 10
part_of located_in
IEA ISO
GO_REF:0000107 (PMID:19490896)
Ensembl RGD
PMID:19490896
GO_REF:0000107
NCBI chr X:104,656,809...104,665,122
Ensembl chr X:104,656,812...104,665,097
G
Psmd2
proteasome 26S subunit ubiquitin receptor, non-ATPase 2
part_of
IBA
GO_REF:0000033
GO_Central
GO_REF:0000033
NCBI chr11:80,248,364...80,258,991
Ensembl chr11:80,248,364...80,259,043
G
Psmd4
proteasome 26S subunit ubiquitin receptor, non-ATPase 4
part_of
IBA
GO_REF:0000033
GO_Central
GO_REF:0000033
NCBI chr 2:182,598,934...182,608,250
Ensembl chr 2:182,598,934...182,608,194
G
Psmd5
proteasome 26S subunit, non-ATPase 5
located_in
ISO
(PMID:19412159), (PMID:19490896)
RGD
PMID:19412159 PMID:19490896
NCBI chr 3:18,133,715...18,151,000
Ensembl chr 3:18,133,717...18,151,029
G
Psmd9
proteasome 26S subunit, non-ATPase 9
located_in
ISO
(PMID:19490896)
RGD
PMID:19490896
NCBI chr12:33,337,182...33,357,468
G
Adrm1
ADRM1 26S proteasome ubiquitin receptor
part_of
IBA
GO_REF:0000033
GO_Central
GO_REF:0000033
NCBI chr 3:167,265,435...167,270,197
Ensembl chr 3:167,265,296...167,270,195
G
Psmd11
proteasome 26S subunit, non-ATPase 11
part_of
IBA
GO_REF:0000033
GO_Central
GO_REF:0000033
NCBI chr10:65,433,853...65,479,774
Ensembl chr10:65,433,853...65,479,766
G
Psmd12
proteasome 26S subunit, non-ATPase 12
part_of
IBA
GO_REF:0000033
GO_Central
GO_REF:0000033
NCBI chr10:92,438,223...92,457,323
Ensembl chr10:92,438,298...92,457,287
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Psmd13
proteasome 26S subunit, non-ATPase 13
part_of
IBA
GO_REF:0000033
GO_Central
GO_REF:0000033
NCBI chr 1:195,964,617...195,976,895
Ensembl chr 1:195,964,138...195,976,905
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Psmd14
proteasome 26S subunit, non-ATPase 14
part_of
ISO IBA
(PMID:22909820) GO_REF:0000033
RGD GO_Central
PMID:22909820
GO_REF:0000033
NCBI chr 3:46,254,338...46,347,076
Ensembl chr 3:46,254,330...46,347,076
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Psmd3
proteasome 26S subunit, non-ATPase 3
part_of
IBA
GO_REF:0000033
GO_Central
GO_REF:0000033
NCBI chr10:83,643,473...83,655,620
Ensembl chr10:83,643,647...83,655,618
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Psmd8
proteasome 26S subunit, non-ATPase 8
part_of
IBA
GO_REF:0000033
GO_Central
GO_REF:0000033
NCBI chr 1:84,474,377...84,481,209
Ensembl chr 1:84,474,378...84,481,209
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Sem1
SEM1 26S proteasome subunit
part_of
IEA
GO_REF:0000104
UniProt
GO_REF:0000104
NCBI chr 4:34,671,848...34,692,005
Ensembl chr 4:34,671,848...34,692,005
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Psma8
proteasome 20S subunit alpha 8
part_of
ISO
(MGI:5494329|PMID:23706739), (MGI:6360105|PMID:31437213), (MGI:6362274|PMID:31358751) (PMID:23706739), (PMID:31358751), (PMID:31437213)
RGD
PMID:23706739 PMID:31358751 PMID:31437213
MGI:5494329 MGI:6360105 MGI:6362274
NCBI chr18:5,863,594...5,928,226
Ensembl chr18:5,863,608...5,928,226
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Psmb10
proteasome 20S subunit beta 10
part_of
ISS ISO
GO_REF:0000024 (MGI:5494329|PMID:23706739) (PMID:23706739)
UniProt RGD
PMID:23706739
GO_REF:0000024 MGI:5494329
NCBI chr19:33,830,958...33,833,442
Ensembl chr19:33,827,229...33,833,626
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Psmb8
proteasome 20S subunit beta 8
part_of
ISS ISO IEA
GO_REF:0000024 (MGI:5494329|PMID:23706739) GO_REF:0000107 (PMID:23706739)
UniProt RGD Ensembl
PMID:23706739
GO_REF:0000024 GO_REF:0000107 MGI:5494329
NCBI chr20:4,652,159...4,655,122
Ensembl chr20:4,652,159...4,655,283
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Psmb9
proteasome 20S subunit beta 9
part_of
ISS ISO
GO_REF:0000024 (MGI:5494329|PMID:23706739) (PMID:23706739)
UniProt RGD
PMID:23706739
GO_REF:0000024 MGI:5494329
NCBI chr20:4,667,044...4,672,512
Ensembl chr20:4,666,046...4,672,512
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Psme4
proteasome activator subunit 4
part_of
ISO IBA
(MGI:5494329|PMID:23706739) GO_REF:0000033 (PMID:23706739)
RGD GO_Central
PMID:23706739
GO_REF:0000033 MGI:5494329
NCBI chr14:104,505,716...104,609,410
Ensembl chr14:104,505,716...104,609,408
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