Protein SMAD4

URN urn:agi-llid:4089
Total Entities 0
Connectivity 1773
Name SMAD4
Description MAD homolog 4 (Drosophila)
Notes transduces signal from TGF-beta; deletion of human homolog gene is associated with pancreatic carcinomas [RGD]

GO Molecular Function transforming growth factor beta receptor, common-partner cytoplasmic mediator activity
protein binding transcription factor activity
RNA polymerase II transcription factor binding transcription factor activity
sequence-specific DNA binding transcription factor activity
chromatin binding
SMAD binding
collagen binding
RNA polymerase II transcription factor binding
filamin binding
protein homodimerization activity
I-SMAD binding
R-SMAD binding
DNA binding
sequence-specific DNA binding
core promoter proximal region sequence-specific DNA binding
transcription regulatory region DNA binding
metal ion binding

GO Cellular Component nucleoplasm
nucleolus
activin responsive factor complex
centrosome
protein complex
transcription factor complex
SMAD protein complex
intracellular
cytoplasm
cytosol
nucleus

GO Biological Process cell proliferation
somite rostral-caudal axis specification
anterior-posterior pattern specification
branching involved in ureteric bud morphogenesis
regulation of transforming growth factor beta2 production
kidney development
metanephric mesenchyme morphogenesis
nephrogenic mesenchyme morphogenesis
atrioventricular canal development
positive regulation of cell proliferation involved in heart valve morphogenesis
epithelial to mesenchymal transition involved in endocardial cushion formation
endocardial cell differentiation
atrioventricular valve formation
cardiac septum development
neural crest cell differentiation
positive regulation of epithelial to mesenchymal transition
sebaceous gland development
regulation of hair follicle development
neuron fate commitment
axon guidance
in utero embryonic development
gastrulation
gastrulation with mouth forming second
brainstem development
tissue morphogenesis
developmental growth
formation of anatomical boundary
SMAD protein signal transduction
intracellular signal transduction
positive regulation of transforming growth factor beta receptor signaling pathway
positive regulation of BMP signaling pathway
positive regulation of SMAD protein import into nucleus
positive regulation of pathway-restricted SMAD protein phosphorylation
regulation of transforming growth factor beta receptor signaling pathway
transforming growth factor beta receptor signaling pathway
BMP signaling pathway
SMAD protein complex assembly
negative regulation of cell growth
endothelial cell activation
negative regulation of cell death
negative regulation of cell proliferation
regulation of cell proliferation
response to transforming growth factor beta
response to hypoxia
positive regulation of transcription, DNA-templated
positive regulation of transcription from RNA polymerase II promoter
negative regulation of transcription, DNA-templated
negative regulation of transcription from RNA polymerase II promoter
regulation of transcription, DNA-templated
regulation of transcription from RNA polymerase II promoter
negative regulation of protein catabolic process
palate development
mesoderm development
endoderm development
transcription, DNA-templated
transcription from RNA polymerase II promoter
transcription initiation from RNA polymerase II promoter
gene expression
regulation of binding

Pathway Cell Cycle
Prostate Cancer Overview
Metastatic Colorectal Cancer Overview
Role of Myostatin in Skeletal Muscles
Myostatin-IGF1 Crosstalk in Skeletal Muscles
Chronic Myeloid Leukemia Overview
Proteins Involved in Pathogenesis of Breast Cancer Related to ERBB2/VEGFR/Akt Signaling Pathway
Proteins Involved in Pathogenesis of Breast Cancer Related to IGF1R/Akt Signaling Pathway
Breast Cancer Related to NOTCH1 Signaling Pathway
Proteins Involved in Pathogenesis of Breast Cancer Related to ESR1 Signaling Pathway
Proteins Involved in Pathogenesis of Breast Cancer Related to NOTCH Signaling Pathway
Basal Breast Cancer
Effects of Hyperthyroidism on Bone Remodeling
Ovarian Cancer Overview
Proteins Involved in Pathogenesis of Prostate Cancer
Activation of Immunoglobulin Class-Switch Recombination
Cancer Overview
Impairment of BMP and TGF-beta Signaling in Familial and Idiopathic Pulmonary Arterial Hypertension
Endothelial Cell Dysfunction in Pulmonary Arterial Hypertension
BMP2 Activates Non-Canonical WNT Signaling in Pulmonary Artery Endothelial Cells
Activation of Glioma Stem Cell Program
Medulloblastoma Overview
Impaired TGFB2 Signaling in Osteoarthritis
Proteins Involved in Pathogenesis of Cataract
Mutations in Neurogenic Transcription Factor Genes in Hirschsprung Disease
STK11 Signaling in Pancreatic Cancer
TGFBR Signaling in Pancreatic Cancer
Pancreatic Ductal Carcinoma
Proteins Involved in Pathogenesis of Pancreatic Cancer
Pancreatic Neuroendocrine Tumors
Proteins Involved in Pathogenesis of Peutz-Jeghers Syndrome
Peutz-Jeghers Syndrome Overview
Growth Factor Signaling in Hepatocellular Carcinoma
NOTCH Signaling in Hepatocellular Carcinoma
Hepatocellular Carcinoma Overview
Vascularization in Hepatocellular Carcinoma
Impaired Osteoblast Function in Gout
Synovial Fibroblast Proliferation in Rheumatoid Arthritis
TGFBR -> MEF/MYOD/MYOG signaling
ActivinR -> SMAD2/3 signaling
TGFBR -> SMAD1/5/9 signaling
ActivinR/BMPR -> SMAD1/5/9 signaling
TGFBR/BMPR -> SMAD2/3 signaling
Dexamethasone-Induced Osteoporosis
Ethanol-Induced Hepatotoxicity
BMP7-ACVR2 Expression Targets
INHBA-ACVR2/ACVR1 Expression Targets
MSTN-ACVR2/ACVR1 Expression Targets
INHBB-ACVR2 Expression Target
BMP2-BMPR2 Expression Targets
BMP4-BMPR2 Expression Targets
BMP15-BMPR2 Expression Targets
BMP7-BMPR2/ACVR2 Expression Targets
GDF5-BMPR2/ACVR2 Expression Targets
BMP6-ACVR2A Expression Targets
MSTN-ACVR2/BMPR Expression Targets
TDGF1-ACVR2B Expression Targets
NODAL-ACVR2B Expression Targets
AMH-AMHR2 Expression Targets
TGFB2-TGFBR1 Expression Targets
TGFB3-TGFBR1 Expression Targets
INHBA-ACVR2/BMPR Expression Targets
TGFB1-TGFBR1 Expression Targets
TGFB1-ACVRL1 Expression Targets
TGFB1-TGFBR1/AP-1 Expression Targets
transforming growth factor-beta Smad dependent signaling pathway
colorectal cancer pathway
pancreatic cancer pathway
transforming growth factor-beta Smad dependent signaling pathway
colorectal cancer pathway
altered transforming growth factor-beta Smad dependent signaling pathway
transforming growth factor-beta Smad dependent signaling pathway

Group SMAD
Tumor suppressors
cell proliferation
somite rostral-caudal axis specification
anterior-posterior pattern specification
branching involved in ureteric bud morphogenesis
regulation of transforming growth factor beta2 production
kidney development
metanephric mesenchyme morphogenesis
nephrogenic mesenchyme morphogenesis
atrioventricular canal development
positive regulation of cell proliferation involved in heart valve morphogenesis
epithelial to mesenchymal transition involved in endocardial cushion formation
endocardial cell differentiation
atrioventricular valve formation
cardiac septum development
neural crest cell differentiation
positive regulation of epithelial to mesenchymal transition
sebaceous gland development
regulation of hair follicle development
neuron fate commitment
axon guidance
in utero embryonic development
gastrulation
gastrulation with mouth forming second
brainstem development
tissue morphogenesis
developmental growth
formation of anatomical boundary
SMAD protein signal transduction
intracellular signal transduction
transforming growth factor beta receptor, common-partner cytoplasmic mediator activity
positive regulation of transforming growth factor beta receptor signaling pathway
positive regulation of BMP signaling pathway
positive regulation of SMAD protein import into nucleus
positive regulation of pathway-restricted SMAD protein phosphorylation
regulation of transforming growth factor beta receptor signaling pathway
transforming growth factor beta receptor signaling pathway
BMP signaling pathway
SMAD protein complex assembly
negative regulation of cell growth
endothelial cell activation
negative regulation of cell death
negative regulation of cell proliferation
regulation of cell proliferation
response to transforming growth factor beta
response to hypoxia
positive regulation of transcription, DNA-templated
positive regulation of transcription from RNA polymerase II promoter
negative regulation of transcription, DNA-templated
negative regulation of transcription from RNA polymerase II promoter
protein binding transcription factor activity
regulation of transcription, DNA-templated
RNA polymerase II transcription factor binding transcription factor activity
sequence-specific DNA binding transcription factor activity
regulation of transcription from RNA polymerase II promoter
negative regulation of protein catabolic process
palate development
mesoderm development
endoderm development
transcription, DNA-templated
transcription from RNA polymerase II promoter
transcription initiation from RNA polymerase II promoter
gene expression
regulation of binding
chromatin binding
SMAD binding
collagen binding
RNA polymerase II transcription factor binding
filamin binding
protein homodimerization activity
I-SMAD binding
R-SMAD binding
DNA binding
sequence-specific DNA binding
core promoter proximal region sequence-specific DNA binding
transcription regulatory region DNA binding
metal ion binding
nucleoplasm
nucleolus
activin responsive factor complex
centrosome
protein complex
transcription factor complex
SMAD protein complex
intracellular
cytoplasm
cytosol
nucleus
Proteins Involved in Pathogenesis of Cowden Syndrome
Genes with Mutations Associated with Pancreatic Cancer
Secreted proteins
Biofluids assayable substances

Source Curated

LocusLink ID 4089
17128
50554
28063

Cell Localization Nucleus
Cytoplasm

GO ID 0003677
0070411
0070412
0001085
0001076
0003682
0005518
0000987
0046872
0000988
0042803
0043565
0003700
0044212
0030616
0030509
0007183
0060395
0036302
0003190
0007411
0003360
0001658
0003279
0008283
0048589
0060956
0007492
0042118
0003198
0048859
0001702
0010467
0001701
0007498
0072133
0060548
0030308
0008285
0042177
0000122
0045892
0072134
0014033
0048663
0060021
0030513
0060391
0003251
0010718
0010862
0045944
0045893
0030511
0051098
0051797
0017015
0032909
0001666
0071559
0048733
0032525
0006367
0006351
0007179
0071141
0032444
0005813
0005737
0005829
0005730
0005654
0005634
0005667
0046332
0031005
0009952
0007369
0035556
0001822
0042127
0006357
0006355
0048729
0006366
0005622
0043234
0010843
0016563
0045941
0001657
0045449
0006350
0005515
0007184
0007182
0009887
0003712
0010003

Alias JIP
DPC4
MADH4
MYHRS
mothers against decapentaplegic homolog 4
MAD homolog 4
SMAD, mothers against DPP homolog 4
deleted in pancreatic carcinoma locus 4
deletion target in pancreatic carcinoma 4
mothers against decapentaplegic, Drosophila, homolog of, 4
AW743858
D18Wsu70e
Smad 4
mothers against DPP homolog 4
deletion target in pancreatic carcinoma 4 homolog
MAD (mothers against decapentaplegic Drosophila) homolog 4
Mothers against decapentaplegic homolog IV
Mothers against decapentaplegic, Drosophila, homolog of, IV
Mothers against DPP homolog IV
OTTHUMP00000163548
OTTMUSP00000023202
OTTMUSP00000023201
SMAD, mothers against DPP homolog IV
deleted in pancreatic carcinoma protein 4
Sma and Mad related protein 4
Smad4 transcription factor
transcription factor DPC4
MAD, mothers against decapentaplegic homolog 4 (Drosophila)
transcription factor Smad4
MAD homolog IV
Deletion target in pancreatic carcinoma IV
DNA segment, Chr 18, Wayne State University 70, expressed
MAD homolog 4 (Drosophila)
homolog of Drosophila Mothers against decapentaplegic,, IV
hSMAD4
MAD (mothers against decapentaplegic homolog 4
MAD (mothers against decapentaplegic homolog IV
MAD (mothers against decapentaplegic, Drosophila) homolog IV
homolog of Drosophila Mothers against decapentaplegic,, 4
MAD, mothers against decapentaplegic homolog 4
SMAD family member 4
Mothers against DPP homolog 4
SMAD4
MAD (mothers against decapentaplegic, Drosophila) homolog 4
Mothers against DPP homolog IV
hSMAD4s

Mouse chromosome position 18 49.51 cM
18 48.0 cM

OMIM ID 600993
260350
174900
175050
139210
603391
187300

Rat chromosome position 18q12.3

Hugo ID 6770

Human chromosome position 18q21.1

Swiss-Prot Accession Q13485
P97471
O70437
A8K405
Q6GTP6
Q9CW56
Q9BYG6
Q811W8

PIR ID S71811

GenBank ID NC_000018
XR_243850
NM_005359
NP_005350
NG_013013
NC_018929
AC_000150
AB043547
BAB40977
ABBA01018682
ABBA01018683
ABBA01018684
AC091551
AF045447
AAC03051
AMYH02034710
AMYH02034711
AMYH02034712
CH471096
EAW62985
EAW62986
EAW62987
AK290770
BAF83459
AU120224
BC002379
AAH02379
BM701399
BX647129
U44378
AAA91041
Q13485
DQ891840
ABM82766
DQ895028
ABM85954
NC_000084
XM_006525702
XP_006525765
XM_006525701
XP_006525764
NM_008540
NP_032566
AC_000040
AAHY01140128
AC132418
AC134447
AY119788
AAM74472
CH466528
EDL09559
EDL09560
EDL09561
AB221585
AK004804
BAB23576
AK170255
AY493561
AAR85497
BC046584
AAH46584
GU072917
ADO32891
GU072918
ADO32892
GU072919
ADO32893
GU072920
ADO32894
U79748
AAB57905
P97471
NC_005117
NM_019275
NP_062148
AC_000086
AABR06095872
AABR06095873
AABR06095874
AAHX01095919
AAHX01095920
AAHX01095921
CH473971
EDM14795
EDM14796
EDM14797
AB010954
BAA83092
AF056002
AAC12781
O70437
NT_010966
NW_001838468
NW_927106
AC_000061
AK303027
BAG64153
Q9BYG6
NT_039674
Q6GTP6
Q811W8
NW_001084736
NW_047516
XM_001001642
BG076431
BG073878
BG086595
XP_001001701
XP_001001668
XM_001001692
XM_001001655
XP_993064
XM_001001701
XP_001001632
XM_924404
XM_924401
XM_924397
XP_001001679
XM_924399
XM_001001668
NW_001030635
XM_987848
AI427147
XP_001001642
XM_987970
XM_987790
XM_001001632
XM_924395
XP_992884
XM_001001679
XM_987755
XM_916691
XP_001001692
XP_001001655
XP_992942
NW_000134
XP_992849

Unigene ID Hs.75862
Mm.100399
Rn.9774

KEGG ID hsa:4089
mmu:17128
rno:50554

Swiss-Prot ID SMAD4_HUMAN
SMAD4_RAT
SMAD4_MOUSE
Q6GTP6_MOUSE
A8K405_HUMAN

Ensembl ID ENSG00000141646
ENSP00000341551
ENST00000342988
ENSMUSG00000024515
ENSMUSP00000025393
ENSMUST00000025393
ENSRNOG00000015634
ENSRNOP00000021113
ENSRNOT00000021113

Homologene ID 31310

Organism Homo sapiens
Mus musculus
Rattus norvegicus

MGI ID 894293
106263

RGD ID 3033

MedScan ID 4089

Primary Cell Localization Nucleus

IPI ID IPI00013404
IPI00198545
IPI00125567
IPI00646338
IPI00644112
IPI00644586
IPI00556778

KEGG pathway Wnt signaling pathway
Cell cycle
TGF-beta signaling pathway
Adherens junction