Protein Map3k4

URN urn:agi-llid:4216
Total Entities 0
Connectivity 126
Name Map3k4
Description mitogen activated protein kinase kinase kinase 4 (predicted)
Notes The central core of each mitogen-activated protein kinase (MAPK) pathway is a conserved cascade of 3 protein kinases: an activated MAPK kinase kinase (MAPKKK) phosphorylates and activates a specific MAPK kinase (MAPKK), which then activates a specific MAPK. While the ERK MAPKs are activated by mitogenic stimulation, the CSBP2 and JNK MAPKs are activated by environmental stresses such as osmotic shock, UV irradiation, wound stress, and inflammatory factors. This gene encodes a MAPKKK, the MEKK4 protein, also called MTK1. This protein contains a protein kinase catalytic domain at the C terminus. The N-terminal nonkinase domain may contain a regulatory domain. Expression of MEKK4 in mammalian cells activated the CSBP2 and JNK MAPK pathways, but not the ERK pathway. In vitro kinase studies indicated that recombinant MEKK4 can specifically phosphorylate and activate PRKMK6 and SERK1, MAPKKs that activate CSBP2 and JNK, respectively but cannot phosphorylate PRKMK1, an MAPKK that activates ERKs. MEKK4 is a major mediator of environmental stresses that activate the CSBP2 MAPK pathway, and a minor mediator of the JNK pathway. Two alternatively spliced transcripts encoding distinct isoforms have been described.

Ariadne Ontology MAP3K

GO Molecular Function nucleotide binding
ATP binding
metal ion binding
transferase activity
transferase activity, transferring phosphorus-containing groups
kinase activity
protein kinase activity
protein serine-threonine kinase activity
MAP kinase kinase kinase activity

GO Cellular Component cytoplasm
perinuclear region of cytoplasm

GO Biological Process activation of MAPKK activity
positive regulation of JUN kinase activity
intracellular protein kinase cascade
MAPK cascade
JNK cascade
chorionic trophoblast cell differentiation
protein phosphorylation
phosphorylation
response to stress
regulation of gene expression
male germ-line sex determination
placenta development
determination of dorsal identity

Pathway c-Jun N-terminal kinases MAPK signaling pathway
c-Jun N-terminal kinases MAPK signaling pathway

Group MAP3K
activation of MAPKK activity
positive regulation of JUN kinase activity
intracellular protein kinase cascade
MAPK cascade
JNK cascade
chorionic trophoblast cell differentiation
protein phosphorylation
phosphorylation
response to stress
regulation of gene expression
male germ-line sex determination
placenta development
determination of dorsal identity
nucleotide binding
ATP binding
metal ion binding
transferase activity
transferase activity, transferring phosphorus-containing groups
kinase activity
protein kinase activity
protein serine-threonine kinase activity
MAP kinase kinase kinase activity
cytoplasm
perinuclear region of cytoplasm

Source Curated
RGD modified
ResNet

MedScan ID 4216
21364

Hugo ID 6856

Human chromosome position 6q26

GenBank ID NC_000006
NM_005922
NP_005913
NM_006724
NP_006715
NC_018917
AC_000138
AL109942
CAC12766
CAI21477
AL139393
CAI20815
CAI20816
AL591045
CAH70639
CAH70640
AL596452
CAI41308
CAI41309
CAI41310
CH471051
EAW47586
EAW47587
AA314177
AF002715
AAB68804
AF116604
AAF71029
AK124430
AK307866
BC136276
AAI36277
BC143735
AAI43736
BC143743
BC143750
BC146770
AAI46771
BQ009477
D86968
BAA13204
Q9Y6R4
NC_000083
NM_011948
NP_036078
AC_000039
AC136921
AF485269
AAM33376
AK014185
BAB29197
AK035823
AK122219
BAC65501
AK133605
BAE21746
AK191363
AK195138
AK199121
AK202372
AK203313
AK204512
BC058719
AAH58719
CK634503
U66240
AAC08286
U85607
AAC53126
U85608
AAC53127
O08648
NC_005100
NM_001107456
NP_001100926
AC_000069
CH474059
EDL83075
NT_025741
NW_001838992
NW_923184
AC_000049
Q9P1M2
NT_039638
NW_001030593
Q3UZV9
Q60671
Q6PDG6
Q80U64
Q8K4X6
Q9CRU1
NW_047553
NW_001084751
NT_007422
Q5VTT9
Q9H408
XM_217804
XP_217804
XM_001063040
XP_001063040
XM_001062918
XP_001062918
XM_001063151
XP_001063151
XM_001063095
XP_001063095
NW_000113
NT_086699
119567971
2352277
27529700
55665553
55665554
56203589
56203590
56204628
56204629
57338495
57338496
6685621
74756814
74761422
74761863
7959712
123792182
12851883
1932803
1932805
20977846
28972095
2992474
37590139
6685609
74227302
81880472
81890126
81911405
81912756
81914756
NC_007307
NW_001495620
XM_589178
XP_589178
AA495283
AA497200
CB333797
CB333798
60688294
82194126
AAH91661
AI943023
AI964926
BC091661
BX510990
BX511188
NC_007124
NW_001510936
Q5BJ15
XM_684744
XP_689836
119567972
148922282
149027485
125833645
126003868
51870475
81684495
Dr.109650
Dr.75420
ZDB-GENE-990603-4
325488
Dr.106221
ZDB-GENE-030131-4213
dre:325488
ZDB-GENE-030131-6687

LocusLink ID 4216
26407
308106
21364
17272
27882
511779
30495
325488
334747
561337
570131

Alias MTK1
MEKK4
MAPKKK4
PRO0412
RP3-473J16.4
mitogen-activated protein kinase kinase kinase 4
MEKK 4
MEK kinase 4
MAP three kinase 1
MAP/ERK kinase kinase 4
MAPK/ERK kinase kinase 4
predicted protein of HQ0412
SSK2/SSK22 MAP kinase kinase kinase, yeast, homolog of
dJ473J16.1 (mitogen-activated protein kinase kinase kinase 4)
Tas
RP17
Mek4b
Rp17a
D17Rp17
D17Rp17e
mKIAA0213
T-associated sex reversal
mitogen activated protein kinase kinase kinase 4
mitogen-activated protein kinase kinase kinase IV
MTK I
OTTHUMP00000017559
OTTHUMP00000017560
MEK kinase IV
SSK2/SSK22 MAP kinase kinase kinase, , homolog of
DNA segment, Chr 17, Roswell Park-17, expressed
RP3-473J16.1
FLJ42439
homolog of yeast SSK2/SSK22 MAP kinase kinase kinase
KIAA0213
MAP three kinase I
MAP/ERK kinase kinase 4 isoform a
MAP/ERK kinase kinase 4 isoform b
MAP/ERK kinase kinase IV
Map3k4
MAPK/ ERK kinase kinase 4
MAPK/ ERK kinase kinase IV
mitogen activated protein kinase kinase kinase 4 (predicted)
cb135
fa02f11
wu:fa02f11
chromosome
fc83c01
scaffold
wu:fc83c01
LOC561337
similar to mitogen-activated protein kinase kinase kinase 4 isoform b
LOC570131

Organism Homo sapiens
Mus musculus
Rattus norvegicus
Bos taurus
Danio rerio

OMIM ID 602425

Unigene ID Hs.390428
Mm.28587
Rn.66100
Mm.386946
Bt.48616
Dr.109650
Dr.75420
Dr.106221

KEGG ID hsa:4216
mmu:26407
rno:308106

Swiss-Prot Accession Q9P1M2
Q9Y6R4
O08648
D3ZGU6
O08649
O70124
Q6PDG6
A6H8W0
B7ZLD3
B9EG75
Q5VTT8
Q5VTT9
Q92612
Q9H408
Q3UZV9
Q60671
Q80U64
Q8K4X6
Q9CRU1

EC Number 2.7.11.25

GO ID 0005524
0004709
0046872
0004672
0007254
0000186
0007243
0019100
0043507
0006950
0005737
0048471
0016301
0000166
0004674
0016740
0016772
0060718
0048263
0016310
0001890
0006468
0010468
0000165
0000287
0003674
0008150
0005575
0004713

PIR ID T03022

Swiss-Prot ID M3K4_MOUSE
M3K4_HUMAN
Q9P1M2_HUMAN
Q6PDG6_MOUSE

Cell Localization Cytoplasm
perinuclear region

IPI ID IPI00623304
IPI00875136
IPI00186536
IPI00386260
IPI00873480
IPI00647525
IPI00119175
IPI00230708
IPI00359194
IPI00870607

Mouse chromosome position 17 8.42 cM
17 6.5 cM

Rat chromosome position 1q11

Homologene ID 31346

MGI ID 1346875
109592
90953

RGD ID 1311192

MedLine Reference 9305639
12466851
11498536
12052864
12878610
14702039
15262978
15601262
15881658
16157600
16256071
7566098
9039502
9079650
9305638
9482735
9827804
9841871
10349636
11042159
11076861
11217851
11991724
12477932
12693553
15044949
15731347
15782199
16141072
16141073
16199873
1665374
1672293
16894626
17145501
1794044
1799792
1845916
2081600
2165966
2272509
2568335
2569442
2574145
2911572
3259947
3455895
3792827
7558022
8111116
8406442
8441630
8835526
9268631
9722950
17015265
10207156
16109975
17496914
17543594

Primary Cell Localization Cytoplasm

Homo sapiens Chromosome position 6q26

Microarray ID 104272_s_at
161568_f_at

FunctionalClass ATP binding
MAP kinase kinase kinase
magnesium ion binding
nucleotide binding
protein serine/threonine kinase
transferase
kinase
metal ion binding
protein kinase
protein-tyrosine kinase

Mus musculus Chromosome position 17 6.5 cM

KEGG pathway MAPK signaling pathway
GnRH signaling pathway

Rattus norvegicus Chromosome position 1q11

Bos taurus Chromosome position 9