Protein Prkca

URN urn:agi-llid:5578
Connectivity 1136
Name Prkca
Description protein kinase C, alpha
Notes A294G mutation of protein kinase C alpha does not detectably affect its biochemical properties in vitro or in vivo, and is at least rare in thyroid neoplasias A294G mutation of protein kinase C alpha does not detectably affect its biochemical properties in vitro or in vivo, and is at least rare in thyroid neoplasias. Activation of PKCalpha stabilizes F-actin and thereby opposes the effect of PKCepsilon on membrane remodeling in T84 cells. Alpha-tocopherol decreases superoxide anion release in human monocytes under hyperglycemic conditions via inhibition of protein kinase C-alpha. Calcium-dependent involucrin expression is inversely regulated by protein kinase C (PKC)alpha and PKCdelta. Calpha activation and phosphorylation of syntaxin 4 and Munc18c are required for the cell surface expression of P-selectin and the consequent binding of neutrophils to endothelial cells. Cis activation of the basal promoter of the PKC alpha gene occurs through an activator protein-2-dependent, phorbol ester-responsive pathway, which suggests an autoregulatory manner of transcription in glioblastoma multiforme cells. Data show that protein kinase Calpha mediates the effect of antiarrhythmic peptide on gap junction conductance. Epidermal and hepatocyte growth factors, but not keratinocyte growth factor, modulate protein kinase Calpha translocation to the plasma membrane. Ets1 serves as an effector for PKCalpha to fulfil certain functions in cancer ce. Filamin A was identified as a direct binding partner of protein kinase Calpha; two binding sites were identified on filamin A; a Ca2+ and phospholipid-dependent association of the regulatory domain of protein kinase C with these sites was revealed. Four residues located near the calcium binding site of PKC alpha are critical for activation and in vivo membrane translocation of the enzyme. Il-2 increased protein kinase C alpha expression, but polysaccharide K decreased it. Inhibition of the oxidative burst, i.e., cellular desensitization, was clearly reversed in cells overexpressing PKC alpha, pointing to PKC alpha as the major transmitter in eliciting the oxidative burst in monocytes/macrophages. Isoforms of protein kinase C and their distribution in human adrenal cortex and tumors. Loss of protein kinase Calpha expression may enhance the tumorigenic potential of Gli1 in basal cell carcinoma. NAG-1 expression is up-regulated by TPA in LNCaP cells through a PKC-dependent pathway involving the activation of NF-kappa B. Only PKC-alpha is involved in the signal transduction cascade leading to neutrophil transepithelial migration. Our findings present original data that PKCalpha is the isoform specifically involved in the proliferation of primary human osteoblasts. PKC alpha expressed in human neutrophils can phosphorylate p47phox and induce both its translocation and NADPH oxidase activation as well as the binding of p47phox to the cytosolic fragment of p22phox. PKC alpha functions as a negative modulator of calcineurin-regulated retinoid X receptor responsive element-dependent transcription during T cell activation. PKC alpha is a critical regulatory element that is required for efficient regulatory volume decrease in HeLa cells. PKC alpha phosphorylates diacylglycerol kinase zeta in cells, and this phosphorylation inhibits its kinase activity to remove cellular diacylglycerol, thereby affecting cell growth. PKC alpha plays a necessary role in mediating calcium-induced differentiation. Failure to regulate PKC alpha in SCC4 carcinoma cells may underlie at least part of the failure of calcium to promote differentiation in these cells. PKC stimulation upregulates neuron-specific ELAV (nELAV) proteins (HuB, HuC, and HuD), and induces nELAV proteins to colocalize with the translocated PKCalpha isozyme on the cytoskeleton, with a concomitant increase of nELAV threonine phosphorylation. PKC-alpha and PKC-epsilon act cooperatively in regulating JNK activation in response to PMA. PKC-alpha-mediated NG2 phosphorylation at Thr(2256) is a key step for initiating cell polarization and motility. PKCA acts upstream of PKCtheta to activate IkappaB kinase and NF-kappaB in T-lymphocytes. PKCA has a major role in cytoskeleton-dependent avidity modulation of ALCAM. PKCalpha C1A-C2 interdomain interactions have a role in lipid-mediated PKCalpha activation. PKCalpha and betaII have roles in the regulation of membrane recycling. PKCalpha is involved in the regulation of Ca2+-induced platelet aggregation. PKCalpha is required for ET-1-induced human myometrial cell growth. PKCalpha is specifically required for TPA-induced ERK(MAPK) signaling to trigger gene expressions of p15(INK4b) and p16(INK4a) leading to HepG2 growth inhibition. PKCalpha may regulate Ets1 activity in invasive breast cancer cells. PKCalpha stimulates NO production in endothelial cells and plays a role in regulation of blood flow in vivo. PKCalpha-S100C/A11-mediated pathway is involved in and essential for the growth inhibition of normal human keratinocytes cells by TGFbeta1. PKD activation plays a central role in NT peptide secretion; upstream regulators of PKD include PKC-alpha and -delta and Rho/ROK. PRKCA-dependent PRKD1 activation modulates ERK signal pathway and endothelial cell proliferation by vascular endothelial growth factor A. Protein kinase C (PKC) is a family of serine- and threonine-specific protein kinases that can be activated by calcium and the second messenger diacylglycerol. PKC family members phosphorylate a wide variety of protein targets and are known to be involved in diverse cellular signaling pathways. PKC family members also serve as major receptors for phorbol esters, a class of tumor promoters. Each member of the PKC family has a specific expression profile and is believed to play a distinct role in cells. The protein encoded by this gene is one of the PKC family members. This kinase has been reported to play roles in many different cellular processes, such as cell adhesion, cell transformation, cell cycle checkpoint, and cell volume control. Knockout studies in mice suggest that this kinase may be a fundamental regulator of cardiac contractility and Ca(2+) handling in myocytes. Protein kinase C alpha negatively regulates cell spreading and motility in breast cancer cells. Protein kinase C alpha plays an important role in activating store-operated Ca2+ channels (SOC) in human mesangial cells. Protein kinase C promotes apoptosis in LNCaP prostate cancer cells through activation of p38 MAPK and inhibition of the Akt survival pathway. REVIEW:Interaction of protein kinase C isozymes with membranes containing anionic phospholipids utilizing fluorescent phorbol esters to probe the properties of the C1 domains. Results demonstrate that a branched signaling pathway involving MEK, ERK, PKCepsilon, PKCalpha, and caveolin-1 regulates collagen expression in normal lung tissue and is perturbed during fibrosis. Results identify targets in RLIP76 for phosphorylation by protein kinase C alpha, which may act as substrates for differential transport of doxorubicin. Results propose a new mechanism by which lithium indirectly inhibits glycogen synthase kinase-3beta via phosphatidylinositol 3 kinase-dependent activation of protein kinase C alpha. Results suggest that the involvement of protein kinase C alpha in carbachol-induced soluble amyloid precursor protein (sAPPalpha) release is negligible, but PKC epsilon may be important in coupling cholinergic receptors with APP metabolism. The data do not support a strong association between single nucleotide polymorphism of PRKCA and spina bifida risk. The novel varepsilon and eta and atypical zeta, but not the conventional alpha and beta and the novel delta PKCs, may be involved in the signaling pathways involved in thrombin-induced human platelet P-selectin expression. There was expression of protein kinase C alpha in abnormal muscle fibers. Protein kinase C isoforms may play a role in the pathogenesis of myofibrillar myopathy. Use of PKC agonists and isozyme-specific pseudosubstrate peptide antagonists suggested a role for PKCalpha and -epsilon in VEGF-mediated DAF up-regulation, mediating A cytoprotective pathway in ECs. We analyzed the dependence of the expression of some selected protein kinase C isoenzymes on the availability and/or action of androgens. Activation required for vitamin C-succinate induced apoptosis of HL-60 cells. Autophosphorylation of PKCalpha limits its sensitivity to DAG; kinase inhibitors augment the DAG sensitivity of PKCalpha by destabilizing the closed conformation. Central role of PKC isoforms and the negative regulatory function of c-Src in the control of stromelysin 3 expression. Colocalizes with virus particles and is required for Respiratory syncytial virus (RSV) fusion to the cell membrane of bronchial epithelial cells. Data demonstrate a pathway of Rho activation involving protein kinase c alpha-dependent phosphorylation of p115Rho guanine exchange factor. Diacylglycerol-responsive PKC isoforms differentially influence CaR agonist-induced release of Ca2+ from internal stores. Greater increase of PKCalpha translocation after PMA treatment in breast cancer erythrocytes compared to controls was observed. In gastric cancer, protein translocation of PLCgamma2 and PKCalpha is critical event in the process of apoptosis induction. Phosphorylation of LRP by PKCalpha modulates the endocytic and signaling function of LRP by modifying its association with adaptor proteins. Protein kinase C alpha associates with phospholipase D1 and enhances basal phospholipase D activity in a protein phosphorylation-independent manner in melanoma cells, which contributes to the cell's high invasive potential. Protein kinase C alpha signaling is activated by apolipoprotein A-I, and it has a role in phosphorylation and stabilization of ATP binding cassette transporter A1 for the high density lipoprotein assembly. Protein kinase C alpha, iota, and theta binding to L-selectin cytoplasmic domain is modulated by receptor phosphorylation. Protein kinase C has a role in chitosan glutamate-mediated tight junction disruption. Protein kinase Calpha transcriptional repression via Sp1 by wild type p53 is involved in inhibition of multidrug resistance 1 P-glycoprotein phosphorylation. Protein phosphorylation and dephosphorylation via PKC and the corresponding protein phosphatases contribute to phosphatidylserine exposure and erythrocyte shrinkage after energy depletion. Role in induction of metallproteinases 1 and 3 in fibroblasts by basic calcium phosphate crystals. The affinity of isolated C1A and C1B domains of PKCalpha and PKCgamma for soluble and membrane-incorporated DAG and phorbol ester were measured by isothermal calorimetry and surface plasmon resonance in order to compare activation mechanisms. The direct PKC-dependent activation of the amyloid protein precursor secretory pathway is compromised by reduced PKCalpha expression and a specific role of this isoform in these mechanisms. The first report of focal adhesion kinase and phosphatidylinositol 3-kinase-dependent PKC-alpha activation in bacterial invasion related to cytoskeletal reorganization. There is a complex interplay between PKCalpha, Syk, and Src involving physical interaction, phosphorylation, translocation within the cell, and functional activity regulation. There is an impairment of degradation of protein kinase c alpha in huntington disease 150q cells that is connected with the sequestration of proteasome on mutant huntingtin aggregates. Two fatty acids inhibited the phorbol 12-myristate 13-acetate (PMA)-induced plasma membrane translocation of protein kinase C (PKC)-alpha and -epsilon. We used publicly available gene expression array data to further understand PKC-a-associated gene expression profiles in NSCLC.

Pathway AdrenergicRb -> STAT3 signaling
vascular endothelial growth factor signaling pathway

GO Molecular Function nucleotide binding
transferase activity
ATP binding
metal ion binding
calcium ion binding
zinc ion binding
kinase activity
protein kinase activity
protein serine-threonine kinase activity
protein kinase C activity
histone kinase activity (H3-T6 specific)
calcium-dependent protein kinase C activity

GO Cellular Component protein complex
membrane
plasma membrane
cytoplasm
membrane raft
cytosol
mitochondrion
nucleus
membrane fraction
synaptosome

GO Biological Process negative regulation of cell proliferation
aging
establishment of protein localization
positive regulation of synaptogenesis
chondrocyte differentiation
central nervous system neuron axonogenesis
negative regulation of heart contraction
regulation of muscle contraction
regulation of the force of heart contraction
learning or memory
positive regulation of exocytosis
inactivation of MAPK activity
negative regulation of insulin receptor signaling pathway
negative regulation of translation
negative regulation of protein amino acid phosphorylation
positive regulation of protein amino acid phosphorylation
regulation of peptidyl-tyrosine phosphorylation
negative regulation of protein kinase activity
positive regulation of smooth muscle cell proliferation
induction of apoptosis by extracellular signals
induction of apoptosis by intracellular signals
regulation of receptor-mediated endocytosis
neutrophil chemotaxis
protein amino acid phosphorylation
peptidyl-threonine phosphorylation
peptidyl-serine phosphorylation
histone H3-T6 phosphorylation
cellular calcium ion homeostasis
response to mechanical stimulus
response to antibiotic
response to toxin
response to reactive oxygen species
response to organic cyclic substance
response to ethanol
response to interleukin-1
response to peptide hormone stimulus
response to estradiol stimulus
response to corticosterone stimulus
induction of positive chemotaxis
positive regulation of inflammatory response
negative regulation of glucose import
intracellular signaling pathway

Ariadne Ontology PKC

Group PKC
negative regulation of cell proliferation
aging
establishment of protein localization
positive regulation of synaptogenesis
chondrocyte differentiation
central nervous system neuron axonogenesis
negative regulation of heart contraction
regulation of muscle contraction
regulation of the force of heart contraction
learning or memory
positive regulation of exocytosis
inactivation of MAPK activity
negative regulation of insulin receptor signaling pathway
negative regulation of translation
negative regulation of protein amino acid phosphorylation
positive regulation of protein amino acid phosphorylation
regulation of peptidyl-tyrosine phosphorylation
negative regulation of protein kinase activity
positive regulation of smooth muscle cell proliferation
induction of apoptosis by extracellular signals
induction of apoptosis by intracellular signals
regulation of receptor-mediated endocytosis
neutrophil chemotaxis
protein amino acid phosphorylation
peptidyl-threonine phosphorylation
peptidyl-serine phosphorylation
histone H3-T6 phosphorylation
cellular calcium ion homeostasis
response to mechanical stimulus
response to antibiotic
response to toxin
response to reactive oxygen species
response to organic cyclic substance
response to ethanol
response to interleukin-1
response to peptide hormone stimulus
response to estradiol stimulus
response to corticosterone stimulus
induction of positive chemotaxis
positive regulation of inflammatory response
negative regulation of glucose import
intracellular signaling pathway
nucleotide binding
transferase activity
ATP binding
metal ion binding
calcium ion binding
zinc ion binding
kinase activity
protein kinase activity
protein serine-threonine kinase activity
protein kinase C activity
histone kinase activity (H3-T6 specific)
calcium-dependent protein kinase C activity
protein complex
membrane
plasma membrane
cytoplasm
membrane raft
cytosol
mitochondrion
nucleus
membrane fraction
synaptosome

MedScan ID 5578

Hugo ID 9393

Human chromosome position 17q22-q23.2

LocusLink ID 5578
18750
24680
146784
276901
268497

Alias AAG6
PKCA
PRKACA
MGC129900
MGC129901
PKC-alpha
protein kinase C alpha type
PKC-A
aging-associated gene 6
AI875142
RP23-145G23.2
OTTMUSP00000003037
protein kinase c-alpha
KPCA
KPCA_HUMAN
LOC146784
PRKCA
protein kinase C alpha
protein kinase C, alpha
Protein kinase C, alpha type (PKC-alpha) (PKC-A)
hypothetical protein LOC146784

Organism Homo sapiens
Mus musculus
Rattus norvegicus

GenBank ID NC_000017
NM_002737
NP_002728
NT_010783
NW_926918
NW_001838454
NG_012206
AC_000060
AC_000149
AC005918
AC005988
AC006263
AC006947
AC009452
AC060796
AF395829
AAK84184
CH471099
EAW89014
CS098008
CAJ00128
AA017296
AB209475
BAD92712
AB451258
BAG70072
AB451383
BAG70197
AF035594
AF035595
AF086287
AK055431
AK125425
AK130736
AL596674
AY633609
AAV33302
BC015855
BC053321
AAH53321
BC062759
BC071767
BC103691
BC107592
BC109273
AAI09274
BC109274
AAI09275
BC122530
BM015483
BQ018414
BU150006
BX648954
CA417000
M22199
AAA60098
X52479
CAA36718
P17252
Q2TSD3
Q59FI5
Q7Z727
EU832768
ACE87779
GQ129311
ACT64445
NC_000077
NM_011101
NP_035231
NT_165773
NW_001030434
AC_000033
AF178928
AAD51851
AL607041
CAM24252
AL645535
CAM15233
CAM15234
AL645963
CAM26857
CH466558
EDL34340
AK047652
AK079091
AK147084
BAE27664
AK163950
BAE37546
BC096493
AAH96493
BF455952
M25811
AAA39934
X52684
CAA36907
X52685
CAA36908
P20444
Q3TQ39
Q4VA93
Q9R0Z6
NC_005109
NM_001105713
NW_001084656
NW_047341
AC_000078
CH473948
EDM06446
BC169007
AAI69007
X07286
CAA30266
P05696
XM_343975
XP_343976
XM_001081588
XP_001081588
NW_000040

OMIM ID 176960

Mouse chromosome position 11 68.0 cM

GO ID 0005524
0004698
0035403
0046872
0000166
0004697
0016740
0008270
0007568
0006874
0021955
0002062
0035408
0000188
0008624
0008629
0050930
0023034
0007611
0008285
0046325
0045822
0046627
0001933
0006469
0017148
0030593
0018105
0018107
0045921
0050729
0001934
0048661
0051965
0006937
0050730
0048259
0002026
0046677
0051412
0032355
0045471
0070555
0009612
0014070
0043434
0000302
0009636
0005737
0005829
0005624
0045121
0005739
0005634
0005886
0043234
0019717
0016301
0004672
0004674
0006468
0016020
0045184
0005509
0007242
0019992
0005515
0007166
0004682
0004713
0004691
0000074

Rat chromosome position 10q32.1

Swiss-Prot Accession B5BU22
P17252
Q7Z727
Q2TSD3
Q59FI5
P20444
Q4VA93
Q3TQ39
Q9R0Z6
P05696
Q15137
Q32M72
Q96RE4

PIR ID S09496
S07104

Unigene ID Hs.531704
Hs.708867
Mm.222178
Rn.207908
Hs.637776
Rn.86669

KEGG ID hsa:5578
mmu:18750
rno:24680

EC Number 2.7.11.13
2.7.11.1
2.7.1.37

Swiss-Prot ID KPCA_HUMAN
KPCA_MOUSE
B5BU22_HUMAN
Q4VA93_MOUSE
Q7Z727_HUMAN

Cell Localization Cytoplasm
Cell membrane

IPI ID IPI00385449
IPI00321446
IPI00884460
IPI00748556
IPI00652220
IPI00201792
IPI00781307
IPI00781159
IPI00780412
IPI00779055
IPI00766874

Homologene ID 55679

RGD ID 3395

MGI ID 97595