Capn11 (calpain 11) - Rat Genome Database

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Gene: Capn11 (calpain 11) Rattus norvegicus
Analyze
Symbol: Capn11
Name: calpain 11
RGD ID: 1302946
Description: Predicted to enable calcium-dependent cysteine-type endopeptidase activity. Predicted to be involved in proteolysis. Predicted to be located in acrosomal vesicle. Predicted to be active in cytoplasm. Orthologous to human CAPN11 (calpain 11); INTERACTS WITH bisphenol A; trichloroethene; 2,3,7,8-tetrachlorodibenzodioxine (ortholog).
Type: protein-coding
RefSeq Status: VALIDATED
Previously known as: calcium-activated neutral proteinase 11; CANP 11
RGD Orthologs
Human
Mouse
Chinchilla
Bonobo
Dog
Squirrel
Pig
Green Monkey
Naked Mole-Rat
Black Rat
Alliance Orthologs
More Info more info ...
Latest Assembly: GRCr8 - GRCr8 Assembly
Position:
Rat AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
GRCr8922,841,500 - 22,863,042 (+)NCBIGRCr8GRCr8GRCr8
GRCr8 Ensembl922,841,522 - 22,863,032 (+)EnsemblGRCr8
mRatBN7.2915,344,059 - 15,365,604 (+)NCBImRatBN7.2mRatBN7.2
mRatBN7.2 Ensembl915,344,089 - 15,365,601 (+)EnsemblmRatBN7.2
UTH_Rnor_SHR_Utx923,930,644 - 23,952,331 (+)NCBIUTH_Rnor_SHR_UtxUTH_Rnor_SHR_Utx
UTH_Rnor_SHRSP_BbbUtx_1.0928,993,324 - 29,015,008 (+)NCBIUTH_Rnor_SHRSP_BbbUtx_1.0UTH_Rnor_SHRSP_BbbUtx_1.0
UTH_Rnor_WKY_Bbb_1.0927,294,084 - 27,315,763 (+)NCBIUTH_Rnor_WKY_Bbb_1.0UTH_Rnor_WKY_Bbb_1.0
Rnor_6.0917,728,845 - 17,750,357 (+)NCBIRnor_6.0Rnor_6.0rn6Rnor6.0
Rnor_6.0 Ensembl917,728,816 - 17,750,363 (+)Ensemblrn6Rnor6.0
Rnor_5.0916,617,234 - 16,638,763 (+)NCBIRnor_5.0Rnor_5.0rn5
RGSC_v3.4910,944,080 - 10,965,592 (+)NCBIRGSC_v3.4RGSC_v3.4rn4
Celera913,088,339 - 13,109,769 (+)NCBICelera
RGSC_v3.1910,946,925 - 10,962,499 (+)NCBI
Cytogenetic Map9q12NCBI
JBrowse:




Gene Ontology Annotations     Click to see Annotation Detail View

Biological Process
proteolysis  (IBA,IEA)

Cellular Component

Molecular Function

References

References - curated
# Reference Title Reference Citation
1. Phylogenetic-based propagation of functional annotations within the Gene Ontology consortium. Gaudet P, etal., Brief Bioinform. 2011 Sep;12(5):449-62. doi: 10.1093/bib/bbr042. Epub 2011 Aug 27.
2. Rat ISS GO annotations from GOA human gene data--August 2006 GOA data from the GO Consortium
3. A genomic analysis of rat proteases and protease inhibitors. Puente XS and Lopez-Otin C, Genome Res 2004 Apr;14(4):609-22.
4. GOA pipeline RGD automated data pipeline
5. Data Import for Chemical-Gene Interactions RGD automated import pipeline for gene-chemical interactions
Additional References at PubMed
PMID:12477932   PMID:16541461  


Genomics

Comparative Map Data
Capn11
(Rattus norvegicus - Norway rat)
Rat AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
GRCr8922,841,500 - 22,863,042 (+)NCBIGRCr8GRCr8GRCr8
GRCr8 Ensembl922,841,522 - 22,863,032 (+)EnsemblGRCr8
mRatBN7.2915,344,059 - 15,365,604 (+)NCBImRatBN7.2mRatBN7.2
mRatBN7.2 Ensembl915,344,089 - 15,365,601 (+)EnsemblmRatBN7.2
UTH_Rnor_SHR_Utx923,930,644 - 23,952,331 (+)NCBIUTH_Rnor_SHR_UtxUTH_Rnor_SHR_Utx
UTH_Rnor_SHRSP_BbbUtx_1.0928,993,324 - 29,015,008 (+)NCBIUTH_Rnor_SHRSP_BbbUtx_1.0UTH_Rnor_SHRSP_BbbUtx_1.0
UTH_Rnor_WKY_Bbb_1.0927,294,084 - 27,315,763 (+)NCBIUTH_Rnor_WKY_Bbb_1.0UTH_Rnor_WKY_Bbb_1.0
Rnor_6.0917,728,845 - 17,750,357 (+)NCBIRnor_6.0Rnor_6.0rn6Rnor6.0
Rnor_6.0 Ensembl917,728,816 - 17,750,363 (+)Ensemblrn6Rnor6.0
Rnor_5.0916,617,234 - 16,638,763 (+)NCBIRnor_5.0Rnor_5.0rn5
RGSC_v3.4910,944,080 - 10,965,592 (+)NCBIRGSC_v3.4RGSC_v3.4rn4
Celera913,088,339 - 13,109,769 (+)NCBICelera
RGSC_v3.1910,946,925 - 10,962,499 (+)NCBI
Cytogenetic Map9q12NCBI
CAPN11
(Homo sapiens - human)
Human AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
GRCh38644,158,820 - 44,184,401 (+)NCBIGRCh38GRCh38hg38GRCh38
GRCh38.p14 Ensembl644,158,811 - 44,184,401 (+)Ensemblhg38GRCh38
GRCh37644,126,557 - 44,152,138 (+)NCBIGRCh37GRCh37hg19GRCh37
Build 36644,234,526 - 44,260,117 (+)NCBIBuild 36Build 36hg18NCBI36
Build 34644,234,525 - 44,260,117NCBI
Celera645,679,690 - 45,705,285 (+)NCBICelera
Cytogenetic Map6p21.1NCBI
HuRef643,847,872 - 43,873,397 (+)NCBIHuRef
CHM1_1644,130,722 - 44,155,885 (+)NCBICHM1_1
T2T-CHM13v2.0643,992,275 - 44,018,561 (+)NCBIT2T-CHM13v2.0
Capn11
(Mus musculus - house mouse)
Mouse AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
GRCm391745,941,117 - 45,970,400 (-)NCBIGRCm39GRCm39mm39
GRCm39 Ensembl1745,941,130 - 45,970,251 (-)EnsemblGRCm39 EnsemblGRCm39
GRCm381745,630,191 - 45,659,709 (-)NCBIGRCm38GRCm38mm10GRCm38
GRCm38.p6 Ensembl1745,630,204 - 45,659,325 (-)Ensemblmm10GRCm38
MGSCv371745,767,153 - 45,796,258 (-)NCBIMGSCv37MGSCv37mm9NCBIm37
MGSCv361745,093,785 - 45,122,890 (-)NCBIMGSCv36mm8
Celera1749,065,616 - 49,089,535 (-)NCBICelera
Cytogenetic Map17CNCBI
cM Map1722.59NCBI
Capn11
(Chinchilla lanigera - long-tailed chinchilla)
Chinchilla AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
ChiLan1.0 EnsemblNW_0049554379,791,021 - 9,804,769 (+)Ensembl
ChiLan1.0NW_0049554379,791,021 - 9,804,769 (+)NCBIChiLan1.0ChiLan1.0
CAPN11
(Pan paniscus - bonobo/pygmy chimpanzee)
Bonobo AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
NHGRI_mPanPan1-v2558,660,714 - 58,691,048 (+)NCBINHGRI_mPanPan1-v2
NHGRI_mPanPan1654,530,809 - 54,561,085 (+)NCBINHGRI_mPanPan1
Mhudiblu_PPA_v0643,753,201 - 43,780,465 (+)NCBIMhudiblu_PPA_v0Mhudiblu_PPA_v0panPan3
PanPan1.1645,040,057 - 45,064,765 (+)NCBIPanPan1.1PanPan1.1panPan2
PanPan1.1 Ensembl645,040,057 - 45,064,765 (+)EnsemblpanPan2panpan1.1
CAPN11
(Canis lupus familiaris - dog)
Dog AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
CanFam3.11212,580,952 - 12,604,973 (+)NCBICanFam3.1CanFam3.1canFam3CanFam3.1
CanFam3.1 Ensembl1212,580,252 - 12,598,633 (+)EnsemblcanFam3CanFam3.1
Dog10K_Boxer_Tasha1212,607,222 - 12,625,764 (+)NCBIDog10K_Boxer_TashaDog10K_Boxer_Tasha
ROS_Cfam_1.01213,063,224 - 13,081,793 (+)NCBIROS_Cfam_1.0ROS_Cfam_1.0
ROS_Cfam_1.0 Ensembl1213,063,290 - 13,081,482 (+)EnsemblROS_Cfam_1.0 Ensembl
UMICH_Zoey_3.11212,588,865 - 12,607,347 (+)NCBIUMICH_Zoey_3.1
UNSW_CanFamBas_1.01212,676,546 - 12,695,304 (+)NCBIUNSW_CanFamBas_1.0
UU_Cfam_GSD_1.01212,770,497 - 12,789,068 (+)NCBIUU_Cfam_GSD_1.0UU_Cfam_GSD_1.0
Capn11
(Ictidomys tridecemlineatus - thirteen-lined ground squirrel)
Squirrel AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
HiC_Itri_2NW_02440494648,052,339 - 48,071,166 (+)NCBIHiC_Itri_2
SpeTri2.0 EnsemblNW_00493647615,889,055 - 15,901,178 (-)EnsemblSpeTri2.0 Ensembl
SpeTri2.0NW_00493647615,889,330 - 15,901,169 (-)NCBISpeTri2.0SpeTri2.0SpeTri2.0
CAPN11
(Sus scrofa - pig)
Pig AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
Sscrofa11.1 Ensembl739,168,991 - 39,191,395 (+)EnsemblsusScr11Sscrofa11.1
Sscrofa11.1739,168,991 - 39,191,934 (+)NCBISscrofa11.1Sscrofa11.1susScr11Sscrofa11.1
Sscrofa10.2744,986,150 - 45,010,543 (+)NCBISscrofa10.2Sscrofa10.2susScr3
CAPN11
(Chlorocebus sabaeus - green monkey)
Green Monkey AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
ChlSab1.11727,980,271 - 28,006,358 (-)NCBIChlSab1.1ChlSab1.1chlSab2
ChlSab1.1 Ensembl1727,980,140 - 27,997,256 (-)EnsemblChlSab1.1 EnsemblchlSab2Vervet-AGM
Vero_WHO_p1.0NW_02366604444,214,982 - 44,241,532 (+)NCBIVero_WHO_p1.0Vero_WHO_p1.0
Capn11
(Heterocephalus glaber - naked mole-rat)
Naked Mole-Rat AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
HetGla 1.0NW_00462475415,673,042 - 15,682,891 (-)NCBIHetGla 1.0HetGla 1.0hetGla2
Capn11
(Rattus rattus - black rat)
Black Rat AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
Rrattus_CSIRO_v1484,233,435 - 84,254,251 (+)NCBIRrattus_CSIRO_v1

Variants

.
Variants in Capn11
107 total Variants
miRNA Target Status (No longer updated)

Predicted Target Of
Summary Value
Count of predictions:306
Count of miRNA genes:161
Interacting mature miRNAs:175
Transcripts:ENSRNOT00000065754, ENSRNOT00000067104
Prediction methods:Microtar, Miranda, Rnahybrid, Targetscan
Result types:miRGate_prediction

The detailed report is available here: Full Report CSV TAB Printer

miRNA Target Status data imported from miRGate (http://mirgate.bioinfo.cnio.es/).
For more information about miRGate, see PMID:25858286 or access the full paper here.


QTLs in Region (GRCr8)
The following QTLs overlap with this region.    Full Report CSV TAB Printer Gviewer
RGD IDSymbolNameLODP ValueTraitSub TraitChrStartStopSpecies
2303170Bp332Blood pressure QTL 3323.730.027arterial blood pressure trait (VT:2000000)mean arterial blood pressure (CMO:0000009)9671488884474983Rat
631211Bw4Body weight QTL45.31retroperitoneal fat pad mass (VT:0010430)retroperitoneal fat pad weight to body weight ratio (CMO:0000635)9534647950346479Rat
2303559Gluco54Glucose level QTL 542blood glucose amount (VT:0000188)blood glucose level (CMO:0000046)9875049253750492Rat
7207814Bmd91Bone mineral density QTL 913.5femur size trait (VT:1000369)femoral neck cross-sectional area (CMO:0001697)9131250552Rat
9589055Scfw5Subcutaneous fat weight QTL 55.550.001subcutaneous adipose mass (VT:1000472)abdominal subcutaneous fat pad weight (CMO:0002069)949664545496645Rat
1641911Alcrsp13Alcohol response QTL 13response to alcohol trait (VT:0010489)brain neurotensin receptor 1 density (CMO:0002068)9621499551214995Rat
1354650Despr5Despair related QTL 54.010.0017locomotor behavior trait (VT:0001392)amount of time spent in voluntary immobility (CMO:0001043)9875049253750492Rat
1300124Cm4Cardiac mass QTL 43.55heart mass (VT:0007028)heart weight to body weight ratio (CMO:0000074)9141763315Rat
61425Cia15Collagen induced arthritis QTL 154.6joint integrity trait (VT:0010548)joint inflammation composite score (CMO:0000919)9534647950416711Rat
631643Bp120Blood pressure QTL 12030.004arterial blood pressure trait (VT:2000000)systolic blood pressure (CMO:0000004)9671488829567695Rat
9589158Gluco65Glucose level QTL 656.820.001blood glucose amount (VT:0000188)plasma glucose level (CMO:0000042)949664545496645Rat
1600365Mcs20Mammary carcinoma susceptibility QTL 203mammary gland integrity trait (VT:0010552)mammary tumor growth rate (CMO:0000344)92103143049025574Rat
7411592Foco8Food consumption QTL 87.40.001eating behavior trait (VT:0001431)feed conversion ratio (CMO:0001312)949664545496645Rat
1298088Edpm11Estrogen-dependent pituitary mass QTL 112.5pituitary gland mass (VT:0010496)pituitary gland wet weight (CMO:0000853)9621499551214995Rat


Expression

RNA-SEQ Expression

alimentary part of gastrointestinal system
appendage
circulatory system
ectoderm
endocrine system
endoderm
exocrine system
hemolymphoid system
hepatobiliary system
integumental system
mesenchyme
mesoderm
musculoskeletal system
nervous system
renal system
reproductive system
respiratory system
sensory system
visual system
8 3 13 67 43 42 25 9 25 6 97 38 5 50 24 61 17 9 9

Sequence


Ensembl Acc Id: ENSRNOT00000065754   ⟹   ENSRNOP00000059934
Type: CODING
Position:
Rat AssemblyChrPosition (strand)Source
GRCr8 Ensembl922,841,522 - 22,863,032 (+)Ensembl
mRatBN7.2 Ensembl915,344,089 - 15,365,601 (+)Ensembl
Rnor_6.0 Ensembl917,728,816 - 17,750,363 (+)Ensembl
Ensembl Acc Id: ENSRNOT00000067104   ⟹   ENSRNOP00000061253
Type: CODING
Position:
Rat AssemblyChrPosition (strand)Source
GRCr8 Ensembl922,841,538 - 22,861,697 (+)Ensembl
mRatBN7.2 Ensembl915,344,089 - 15,364,253 (+)Ensembl
Rnor_6.0 Ensembl917,734,345 - 17,749,942 (+)Ensembl
Ensembl Acc Id: ENSRNOT00000114818   ⟹   ENSRNOP00000083385
Type: CODING
Position:
Rat AssemblyChrPosition (strand)Source
GRCr8 Ensembl922,841,522 - 22,863,032 (+)Ensembl
mRatBN7.2 Ensembl915,344,089 - 15,365,601 (+)Ensembl
RefSeq Acc Id: NM_001002806   ⟹   NP_001002806
RefSeq Status: VALIDATED
Type: CODING
Position:
Rat AssemblyChrPosition (strand)Source
GRCr8922,841,530 - 22,863,042 (+)NCBI
mRatBN7.2915,344,089 - 15,365,604 (+)NCBI
Rnor_6.0917,728,845 - 17,750,357 (+)NCBI
Rnor_5.0916,617,234 - 16,638,763 (+)NCBI
RGSC_v3.4910,944,080 - 10,965,592 (+)RGD
Celera913,088,339 - 13,109,769 (+)RGD
Sequence:
RefSeq Acc Id: XM_039084029   ⟹   XP_038939957
Type: CODING
Position:
Rat AssemblyChrPosition (strand)Source
GRCr8922,841,500 - 22,863,042 (+)NCBI
mRatBN7.2915,344,059 - 15,365,500 (+)NCBI
RefSeq Acc Id: XM_063267561   ⟹   XP_063123631
Type: CODING
Position:
Rat AssemblyChrPosition (strand)Source
GRCr8922,841,500 - 22,861,702 (+)NCBI
RefSeq Acc Id: XM_063267562   ⟹   XP_063123632
Type: CODING
Position:
Rat AssemblyChrPosition (strand)Source
GRCr8922,841,500 - 22,863,042 (+)NCBI
RefSeq Acc Id: XM_063267563   ⟹   XP_063123633
Type: CODING
Position:
Rat AssemblyChrPosition (strand)Source
GRCr8922,841,500 - 22,862,174 (+)NCBI
RefSeq Acc Id: XM_063267564   ⟹   XP_063123634
Type: CODING
Position:
Rat AssemblyChrPosition (strand)Source
GRCr8922,841,500 - 22,859,102 (+)NCBI
RefSeq Acc Id: NP_001002806   ⟸   NM_001002806
- UniProtKB: Q6IE72 (UniProtKB/Swiss-Prot),   Q4V8Q1 (UniProtKB/Swiss-Prot),   D3ZXF4 (UniProtKB/TrEMBL)
- Sequence:
Ensembl Acc Id: ENSRNOP00000061253   ⟸   ENSRNOT00000067104
Ensembl Acc Id: ENSRNOP00000059934   ⟸   ENSRNOT00000065754
RefSeq Acc Id: XP_038939957   ⟸   XM_039084029
- Peptide Label: isoform X1
- UniProtKB: Q6IE72 (UniProtKB/Swiss-Prot),   Q4V8Q1 (UniProtKB/Swiss-Prot)
Ensembl Acc Id: ENSRNOP00000083385   ⟸   ENSRNOT00000114818
RefSeq Acc Id: XP_063123632   ⟸   XM_063267562
- Peptide Label: isoform X3
- UniProtKB: Q6IE72 (UniProtKB/Swiss-Prot),   Q4V8Q1 (UniProtKB/Swiss-Prot)
RefSeq Acc Id: XP_063123633   ⟸   XM_063267563
- Peptide Label: isoform X4
- UniProtKB: F1M7U2 (UniProtKB/TrEMBL)
RefSeq Acc Id: XP_063123631   ⟸   XM_063267561
- Peptide Label: isoform X2
- UniProtKB: F1M7U2 (UniProtKB/TrEMBL)
RefSeq Acc Id: XP_063123634   ⟸   XM_063267564
- Peptide Label: isoform X5
- UniProtKB: A6JIX3 (UniProtKB/TrEMBL)
Protein Domains
Calpain catalytic   EF-hand

Protein Structures
Name Modeler Protein Id AA Range Protein Structure
AF-Q4V8Q1-F1-model_v2 AlphaFold Q4V8Q1 1-716 view protein structure

Transcriptome

eQTL   View at Phenogen
WGCNA   View at Phenogen
Tissue/Strain Expression   View at Phenogen

Promoters
RGD ID:13696538
Promoter ID:EPDNEW_R7063
Type:multiple initiation site
Name:Capn11_1
Description:calpain 11
SO ACC ID:SO:0000170
Source:EPDNEW (Eukaryotic Promoter Database, http://epd.vital-it.ch/)
Experiment Methods:Single-end sequencing.
Position:
Rat AssemblyChrPosition (strand)Source
Rnor_6.0917,728,843 - 17,728,903EPDNEW

Additional Information

Database Acc Id Source(s)
AGR Gene RGD:1302946 AgrOrtholog
BioCyc Gene G2FUF-28180 BioCyc
Ensembl Genes ENSRNOG00000026025 Ensembl, ENTREZGENE
Ensembl Transcript ENSRNOT00000065754 ENTREZGENE
  ENSRNOT00000067104 ENTREZGENE
  ENSRNOT00000114818 ENTREZGENE
Gene3D-CATH 2.60.120.380 UniProtKB/Swiss-Prot
  Cysteine proteinases UniProtKB/Swiss-Prot
  EF-hand UniProtKB/Swiss-Prot
IMAGE_CLONE IMAGE:7457866 IMAGE-MGC_LOAD
InterPro C2_III UniProtKB/Swiss-Prot
  Calpain_cysteine_protease UniProtKB/Swiss-Prot
  Calpain_domain_III UniProtKB/Swiss-Prot
  Calpain_III UniProtKB/Swiss-Prot
  Calpain_III_sf UniProtKB/Swiss-Prot
  EF-hand-dom_pair UniProtKB/Swiss-Prot
  EF_Hand_1_Ca_BS UniProtKB/Swiss-Prot
  EF_hand_dom UniProtKB/Swiss-Prot
  Papain-like_cys_pep_sf UniProtKB/Swiss-Prot
  Pept_cys_AS UniProtKB/Swiss-Prot
  Peptidase_C2_calpain_cat UniProtKB/Swiss-Prot
KEGG Report rno:408218 UniProtKB/Swiss-Prot
MGC_CLONE MGC:114218 IMAGE-MGC_LOAD
NCBI Gene 408218 ENTREZGENE
PANTHER CALPAIN UniProtKB/Swiss-Prot
  CALPAIN-11 UniProtKB/Swiss-Prot
Pfam Calpain_III UniProtKB/Swiss-Prot
  Peptidase_C2 UniProtKB/Swiss-Prot
PhenoGen Capn11 PhenoGen
PRINTS CALPAIN UniProtKB/Swiss-Prot
PROSITE CALPAIN_CAT UniProtKB/Swiss-Prot
  EF_HAND_1 UniProtKB/Swiss-Prot
  EF_HAND_2 UniProtKB/Swiss-Prot
  THIOL_PROTEASE_CYS UniProtKB/Swiss-Prot
RatGTEx ENSRNOG00000026025 RatGTEx
SMART calpain_III UniProtKB/Swiss-Prot
  CysPc UniProtKB/Swiss-Prot
  EFh UniProtKB/Swiss-Prot
Superfamily-SCOP SSF47473 UniProtKB/Swiss-Prot
  SSF49758 UniProtKB/Swiss-Prot
  SSF54001 UniProtKB/Swiss-Prot
UniProt A0A8I5ZX40_RAT UniProtKB/TrEMBL
  A6JIX3 ENTREZGENE, UniProtKB/TrEMBL
  CAN11_RAT UniProtKB/Swiss-Prot
  D3ZXF4 ENTREZGENE, UniProtKB/TrEMBL
  F1M7U2 ENTREZGENE, UniProtKB/TrEMBL
  Q4V8Q1 ENTREZGENE
  Q6IE72 ENTREZGENE
UniProt Secondary Q6IE72 UniProtKB/Swiss-Prot


Nomenclature History
Date Current Symbol Current Name Previous Symbol Previous Name Description Reference Status
2005-02-14 Capn11  calpain 11      Symbol and Name status set to provisional 70820 PROVISIONAL